ORDER CANUELLOIDA (POLYARTHRA according by He et al.(2023)


CANUELLIDAE Lang, 1944

COULLANA Por, 1984

  • Coullana canadensis (Willey, 1923) [north America; marine]
  • Canuella elongata Wilson C.B., 1932 (synonymy of Scottolana canadensis (Willey, 1923)

    ECHINOSUNARISTES Huys, 1995

  • Echinosunaristes bathyalis Huys, 1995 [Atlantic Ocean]


    ELANELLA Por, 1984

  • Elanella elantica (Por, 1967) [Red Sea]
  • Elanella haifensis Drumm et al., 2015 [Israel: marine]
  • Elanella paenelanitica (Fiers, 1982) [New Guinea; marine]


    GALAPACANUELLA Mielke, 1979

  • Galapacanuella beckeri Mielke, 1979 [Galapagos; marine]


    IFANELLA Vervoort, 1964

  • Ifanella chacei Vervoort, 1964 [Carolina Islands; marine]


    INTERCANUELLA Becker & Schriever, 1979

  • Intercanuella lima Becker & Schriever, 1979 [questionable taxon (Por, 1984)]


    INTERSUNARISTES Huys, 1995

  • Intersunaristes curticaudatus (Thompson & A. Scott, 1903) [Sri Lanka; ?]
  • Intersunaristes dardani (Humes & Ho, 1969) [Western Indian Ocean]


    MICROCANUELLA Mielke, 1994

  • Microcanuella bisetosa Mielke, 1994 [Costa Rica; marine]
  • Microcanuella secunda Richter, 2015 [Great Meteor Seamount]


    NATHANIELLA Por, 1984

  • Nathaniella reichi Por, 1984 [Israel?:?]


    PARASUNARISTES Fiers, 1982

  • Parasunaristes chelicerata (Por & Marcus, 1972) [Suez canal; marine]
  • Parasunaristes cucullaris Fiers, 1982 [New Guinea; marine]


    SCOTTOLANA Huys, 2009

    "The genus Scottolana Huys, 2009 was proposed by Por (1967) to reallocate three species, Sunaristes inopinata Thompson and Scott, 1903, Sunaristes longipes Thompson and Scott, 1903, and Sunaristes curticaudata Thompson and Scott, 1903. However, the genus name was unavailable because Por (1967) did not designate a type species and his generic diagnosis was not informative to express differential morphological characteristics from other genera (Huys 2009). Huys (2009) proposed and re-established the generic name as Scottolana Huys, 2009 in accordance with ICZN art. 16.1, with a new type species, Scottolana geei Mu and Huys, 2004 (ICZN Art. 13.3). To date, the genus Scottolana accommodates 16 species" (Song et al., 2018 [Thailand; marine]

  • Scottolana antillensis Fiers, 1984 [West Indian Islands; marine]
  • Scottolana brevifurca (Wells, 1967)
  • Scottolana bulbifera (Chislenko, 1971) [Japan; marine]
  • Scottolana bulbosa (Por, 1964)
  • Scottolana canadensis (Willey, 1923) accepted as Coullana canadensis (Willey, 1923) accepted as Canuella canadensis Willey, 1923
  • Scottolana curticaudata (Thompson I.C. & Scott A., 1903) accepted as Intersunaristes curticaudatus (Thompson I.C. & Scott A., 1903)
  • Scottolana daecheonensis Bang et al. 2022 [Korea] (Daecheonensis in original description)
  • Scottolana dissimilis Fiers, 1982 [New Guinea; marine]
  • Scottolana geei Fang-Hong Mu & Huys, 2005 [China; marine]
  • Scottolana gomezi Nazari et al., 2018
  • Scottolana glabra Fiers, 1982 [New Guinea; marine]
  • Scottolana huysi Song et al., 2018 [Thailand; marine]
  • Scottolana inopinata (Thompson I.C. & Scott A., 1903)
  • Scottolana jasani Song et al., 2018 [Korea; marine]
  • Scottolana longipes (Thompson & A. Scott, 1903) [Europe; marine]
  • Scottolana oleosa Wells & Rao, 1987 [Andaman islands; marine]
  • Scottolana rostrata Wells & Rao, 1987 [Andaman islands; marine]
  • Scottolana scotti (Sewell, 1940))
  • Scottolana tumidiseta Wells & Rao, 1987 [Andaman islands; marine]
  • Scottolana uxoris Por, 1983 [Red Sea]
  • Scottolana wonchoeli Bang et al. 2022 [Korea] (Wonchoeli in original description)


    LONGIPEDIIDAE Sars, 1903 sensu Lang, 1944

    The family Longipediidae was initially established by Boeck [15] under the name “Afdeling Longipedina” as the primary group of Harpacticidae. However, several authors [2,16,17] mistakenly attributed the family Longipediidae to Sars, 1903, causing confusion. However these inaccuracies were pointed out and corrected by Wells Khodami et al. proposed a new order Canuelloida Khodami, MacArthur, BlancoBercial and Martinez Arbizu, 2017, including Canuellidae Lang, 1944 and Longipediidae Boeck, 1865; however, this article was retracted by the authors in 2020, due to inaccuracies in data and problems with data processing. Although the World of Copepods database still shows the classification of the order Canuelloida based on the withdrawn paper, the family Longipediidae should be returned to belonging to the order Harpacticoida" (from: Bang et al., 2021)


    Longipediinae Boeck, 1865

    LONGIPEDIA Claus, 1863

    "The taxonomic structure of the genus Longipedia Claus, 1863 has undergone drastic changes since its creation to accommodate the type species L. coronata Claus, 1863, collected in Germany (Helgoland) and Italy (Naples), and poorly described by Claus in 1863 (for revisions see Lang, 1948: 152–155, and Wells, 1980). In his revision, Wells (1980) recognized 10 valid species within Longipedia, four species incertae sedis, and one species nomen dubium. Since then, two valid species (Longipedia andamanica nipponica Itô, 1985, and Longipedia spinulosa Itô, 1981), and one species incertae sedis (Longipedia sp. Fiers, 1984) have been added" (from Gomez, 2001). Recently Schizas et al. (2015) described a new taxon, L. gonzalezi from Puerto Rico, and in the same paper they discuss the phylogenetic position of the Polyarthra (a taxon consisting of the families Longipediidae and Canuellidae, whose affinities with Harpacticoida have been questioned)


  • Longipedia americana Wells, 1980 *
  • Longipedia andamanica Wells, 1980
  • Longipedia andamanica nipponica Ito, 1985
  • Longipedia australica Nicholls, 1941
  • Longipedia brevispinosa Wells, 1980
  • Longipedia coronata Claus, 1863
  • Longipedia corteziensis Gómez, 2001 *
  • Longipedia ferox Krichagin, 1877
  • Longipedia gonzalezi Schizas et al. 2015 [Puerto Rico;mesophotic coral reefs]
  • Longipedia helgolandica Wells, 1980 *
  • Longipedia kikuchii Wells & Rao. 1987 [Andaman and Nicobar Islands]
  • Longipedia koreana Bang et al., 2021 [Korea]
  • Longipedia longispina Monard, 1928
  • Longipedia minor Lang, 1948
  • Longipedia mourei Wells, 1980
  • Longipedia nichollsi Wells, 1980
  • Longipedia paguri Müller W., 1884
  • Longipedia spinulosa Ito, 1981
  • Longipedia pirgos Wells, 1980
  • Longipedia pontica Wells, 1980
  • Longipedia rosea Lang, 1948 (nomen dubium)
  • Longipedia santacruzensis (Mielke, 1979) *
  • Longipedia scotti Lang, 1948
  • Longipedia thailandensis Chullasorn et al., 2008 [Thailand; brackish waters]
  • Longipedia ulleungensis Bang et al., 2021 [Korea]
  • Longipedia weberi Lang, 1948

    * helgolandica species-group, according to Gomez (2001)

    KEY TO SPECIES OF LONGIPEDIIDAE



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