 CLETOPSYLLIDAE Huys & Lee, 1998/99 
"The family Cletopsyllidae includes six genera and 14 species, which are rare
in marine benthic habitats (Boxshall and Halsey 2004). However, Cletopsyllidae does
have a broad global distribution and has been found in the Indian Ocean (Retrocalcar
secundus (Nicholls, 1945); Cletopsyllus bacescui Marcus, 1976; Bathycletopsyllus hexarthra
Huys & Lee, 1999), the Pacific Ocean (R. sagamiensis (Itô, 1971); Isocletopsyllus
maximus Song, Kim & Hwang, 2010), the Atlantic Ocean and the Caribbean Sea (C.
papillifer Willey, 1935; R. brattstroemi (Geddes, 1981); C. rotundifera Fiers, 1986),
and the Mediterranean Sea (I. tertius (Por, 1964); I. quartus (Soyer, 1966); I. sardus
Addis, Floris & Carcupino, 2011)" (from: Woo-Bang et al., 2014)
CLETOPSYLLUS s.l. Willey, 1935
Huys & Lee (1998/99) splitted the genus Cletopsyllus (formerly in the family Normanellidae Lang, 1944, subfamily Cletopsyllinae Huys & Willems, 1989) in five genera: Cletopsyllus s. str. Willey, 1935 ; Retrocalcar Huys & Lee, 1998/99; Bathycletopsyllus Huys & Lee, 1998/99; Isocletopsyllus Huys & Lee, 1998/99; Pentacletopsyllus Woo-Bang et al., 2014.
CLETOPSYLLUS sensu Huys & Lee, 1998/99
Type-species: Cletopsyllus papillifer Willey, 1935
Cletopsyllus bacescui Marcus, 1976 [Indian Ocean]
Cletopsyllus brattstroemi Geddes, 2011 [Bahamas]
Cletopsyllus papillifer Willey, 1935 [Bermuda; marine]
Cletopsyllus quartus Marcus 1976 [Indian Ocean] (=Isocletopsyllus quartus (Soyer, 1966)
Cletopsyllus rotundifera Fiers, 1986 [West Indian Islands; marine]
Cletopsyllus sagamiensis Itô Tat, 1971 accepted as Retrocalcar sagamiensis (Itô Tat, 1971)
Cletopsyllus secundus Nicholls, 1945 accepted as Retrocalcar secundus (Nicholls, 1945)
Cletopsyllus tertius Por, 1964 accepted as Isocletopsyllus tertius (Por, 1964)
RETROCALCAR Huys & Lee, 1998/99
Type-species: Cletopsyllus brattstroemi Geddes, 1981 = Retrocalcar brattstroemi (Geddes, 1981)
Retrocalcar brattstroemi (Geddes, 1981) [Bahamas; marine]
Retrocalcar secundus (Nicholls, 1945) [West Australia; marine]
Retrocalcar sagamiensis (Ito, 1971) [Sagami Bay; marine]
BATHYCLETOPSYLLUS Huys & Lee, 1998/99
Type-species: Bathycletopsyllus hexarthra Huys & Lee, 1998/99
Bathycletopsyllus hexarthra Huys & Lee, 1998/99 [Indian Ocean, La Réunion; marine sand]
ISOCLETOPSYLLUS Huys & Lee, 1998/99
Type-species: Cletopsyllus tertius Por, 964 = Isocletopsyllus tertius (Por, 1964)
Isocletopsyllus maximus Song et al., [Korea; sandy bottoms]
Isocletopsyllus quartus (Soyer, 1966) [France; marine] (species inquirenda)
Isocletopsyllus sardus Adis et al., 2011
Isocletopsyllus tertius (Por, 1964) [Israel; marine] (species inquirenda)
PENTACLETOPSYLLUS Woo-Bang et al., 2014
Pentacletopsyllus montagni Woo-Bang et al., 2014 [Mexico; marine]
GENUS AND SPECIES INQUIRENDA : Pseudocletopsyllus Vervoort, 1964 and Pseudocletopsyllus spiniger Vervoort 1964
RELEVANT LITERATURE
Huys R. & W. Lee, 1998/99. On the relationships of the Normanellidae and the recognition of Cletopsyllidae grad. nov. (Copepoda, Harpacticoida). Zool. Anz, 237: 267-290
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