LAOPHONTOPSIDAE Huys & Willems, 1989
LAOPHONTOPSIS Sars, 1908 (syn. Cleta Claus, 1863)
Laophontopsis borealis Huys & Willems, 1989 [Europe; north Africa, marine]
Laophontopsis brevirostris (Claus, 1863) (= Paralaophonte brevirostris (Claus, 1863)
Laophontopsis forcipata (Claus, 1866)
Laophontopsis lamellifera (Claus, 1863) [Europe; north Africa, marine]
Laophontopsis lilljeborgia (Krichagin, 1877)
Laophontopsis minuticornis (Buchholz, 1874)
Laophontopsis monardi Huys & Willems, 1989 [Europe; north Africa, marine]
Laophontopsis parvula (Claus, 1866)
Laophontopsis secunda (Sewell, 1924) (syn, Telodocus secundus (Sewell, 1924)
Laophontopsis serrata (Claus, 1863) (syn. Laophonte serrata (Claus, 1863)
Laophontopsis setigera (Krichagin, 1877)
Laophontopsis similis (Krichagin, 1873) (syn. Heterolaophonte similis fortior (Krichagin, 1873)
Laophontopsis thalestris (Krichagin, 1877)
Laophontopsis uncinata (Czerniavski, 1868) (syn. Heterolaophonte uncinata (Czerniavski, 1868)
ACULEOPSIS Huys & Willems, 1989
Aculeopsis longisetosa Huys & Willems, 1989
TELODOCUS Huys & Willems, 1989
Telodocus secundus Huys & Willems, 1989
THOMPSONULIDAE Lang, 1944 sensu Huys & Gee, 1990
Huys & Gee (1990) raised to family rank the subfamily Thompsonulinae Lang, 1944, formerly included in the family Tachidiidae (Lang, 1948).
The family presently includes two genera, viz. Thompsonula T. Scott, 1905 and Caribbula Huys & Gee, 1990.
THOMPSONULA T. Scott, 1905 (syn. Echinocornus Wilson C.B., 1932; Rathbunula Wilson C.B., 1932)
Thompsonula agilis (Wilson C.B., 1932)
Thompsonula curticaudata (C.B. Wilson, 1932) [USA; marine sand]
Thompsonula hyaenae (I.C. Thompson, 1889) [Europe, USA, Africa; marine]
Thompsonula hyaenae elongata Gee, 1988 (syn. Caribbula elongata (Gee, 1988)
Thompsonula pectinata (Wilson C.B., 1932)
CARIBBULA Huys & Gee, 1990
Caribbula elongata (Gee, 1988) [Mexico; marine coastal sand]
Caribbula fleegeri Huys & Gee, 1990 [USA; marine, continental shelf]
RELEVANT LITERATURE
Huys R. & J.M. Gee. 1990. A revision of Thompsonulidae Lang, 1944 (Copepoda: Harpacticoida). Zool. J. Linnean Soc., 99: 1-49.
ZOSIMEIDAE Seifried, 2003
"The family Zosimeidae, currently composed of the genera Zosime Boeck, 1873, Peresime Dinet,
1974, Heterozosime Kim &Lee. 2021, Acritozosime Kim & Lee, 2021 and Pseudozosime T. Scott, 1912, was erected by Seifried (2003) based on unique apomorphies in the mouth
parts, P1 endopod, and P5 (Seifried 2003; Huys & Clark 2009). These genera can easily be distinguished from each
other by the segmentation of the endopods of P2-P4 (Wells 2007; Koller & George 2011). Although the family is
small, presently accommodating only 19 species/subspecies and mainly from sublittoral marine habitats (Wells
2007; Koller & George 2011), many unnamed species have been reported in the literature and are awaiting
description (Carey & Montagna 1982; Mu et al. 2002; Seifried 2003; Chertoprud et al. 2009; Huys & Clark 2009).
Zosime is the most speciose genus in the family, currently comprising 16 species/subspecies and assuming a wide
geographical distribution (Koller & George 2011). Recently, Koller & George (2011) noted that some species
display derived features in the antennule, fifth leg, and caudal ramus, and suggested that a phylogenetic study based
on the type material of most species is necessary to confirm or reject the monophyletic status of the genus" (accordin to to Huys & Clark, 2016)
ZOSIME Boeck, 1873"Based on morphological comparison of female character states, the genus Zosime can be subdivided
into three provisional groups, anneae-, incrassata-, and typica-group. These groups are distinguished from each other by
differences in the antennule, antennary exopod, mandibular rami, armature of swimming legs, fifth leg, and caudal rami" (Kim et al., 2016)
Zosime
anneae Koller & George, 2011 [Atlantic Ocean]
Zosime atlanticola Bodin, 1968 (?)
Zosime bathybia Bodin, 1968 (?)
Zosime bathyalis Por, 1967
Zosime bergensis Drzycimski, 1968
Zosime carsteni Pointner, 2017
Zosime changi Kim & Lee, 2019
Zosime comata Kim & Lee, 2019
Zosime destituta Kim et al., 2016 [Korea]
Zosime eliasi Pointner, 2017
Zosime erythraea Por, 1967
Zosime fusiformis Brady, 1880 (syn. Jonesiella fusiformis (Brady & Robertson, 1876)
Zosime gisleni Lang, 1948
Zosime gymnokosmosa Kim & Lee, 2019
Zosime incrassata incrassata Sars, 1910
Zosime incrassata bathybia Bodin, 1968
Zosime major Sarts, 1919
Zosime mediterranea Lang, 1948
Zosime montagnai Yeom et al., 2022 [Mexico; marine]
Zosime mexicana Gomez & Rivera, 2023 [USA]
Zosime pacifica Fiers, 1991
Zosime paramajor Bodin, 1968
Zosime paratypica Becker & Schriever, 1979
Zosime reducta Becker & Schriever, 1979 (syn. Peresime reducta (Becker & Schriever, 1979)
Zosime reyssi Dinet, 1974
Zosime spinulosa Brady & Robertson, 1876 (syn, Danielssenia typica Boeck, 1873)
Zosime thistlei Yeom et al., 2022 [Mexico; marine]
Zosime tunnelli Yeom et al., 2022 [Mexico; marine]
Zosime typica Boeck, 1873
Zosime valida Sars, 1919
Zosime xiximi Gomez & Rivera, 2023 [USA]
SPECIES INCERTAE SEDIS OR INQUIRENDAE
Zomine rubra Thompson, 1888 (nomen nudum)
HETEROZOSIME Kim & Lee, 2021 [Indian Ocean] Heterozosime klemie Gomez & Rivera, 2023 [USA]
Heterozosime tenuis Kim & Lee, 2021 [Indian Ocean]
ACRITOZOSIME Kim & Lee, 2021 [Indian Ocean]
Acritozosime sigsbeeiensis Gomez & Rivera, 2023 [USA]
Acritozosime spinesco Kim & Lee, 2021 [Indian Ocean]
PSEUDOZOSIME T. Scott, 1912
Pseudozosime brownei T. Scott, 1912
PERESIME Dinet, 1974
Peresime abyssalis Dinet, 1974
Peresime pryocta Gomez & Rivera, 2023 [USA]
Peresime pryorae Yeom et al., 2022 [Mexico; marine]
Peresime reducta (Becker & Schriever, 1979)
PORCELLIDIIDAE Boeck, 1865
BREVIFRONS Harris V.A., 1994
Brevifrons faviolatum Harris V.A., 1994
CEREUDORSUM Harris V.A., 2014
Cereudorsum verrucosum Harris V.A., 2014
CLAVIGOFERA Harris V.A. & Iwasaki, 1996
Clavigofera clavigera (Pesta, 1935)
Clavigofera echinophila (Humes & Gelerman, 1962)
Clavigofera laurencia (Hicks, 1982)
Clavigofera pacifica Harris V.A. & Iwasaki, 1996
Clavigofera ulva (Hicks, 1982)
CLUNIA Harris V.A., 2014
Clunia cocosensis Harris V.A., 2014
DILATATIOCAUDA Harris V.A., 2002
Dilatatiocauda bipartita (Kim I.H. & H.S. Kim, 1997)
Dilatatiocauda dilatata (Hicks, 1971)
Dilatatiocauda medialis Harris V.A., 2002
Dilatatiocauda multidenticulata Harris V.A., 2002
Dilatatiocauda plana (Tiemann, 1977)
Dilatatiocauda retroseta Harris, 2002 [Australia]
Dilatatiocauda tristanensis (Wiborg, 1964) KEY TO SPECIES OF DILATATIOCAUDA
GEDDESIA Harris V.A., 2014
Geddesia quadrata Harris V.A., 2014
Geddesia trisetosa (Geddes, 1968)
KENSAKIA Harris V.A. & Iwasaki, 1997
Kensakia acuta (Kim I.H. & Kim H.S., 1997)
Kensakia acuticaudata (Thompson I.C. & Scott A., 1903)
Kensakia aiiroa Harris V.A. & Iwasaki, 1997
Kensakia australis Harris V.A., 2014
Kensakia parva Harris V.A. & Iwasaki, 2009
Kensakia shimodensis Harris V.A. & Iwasaki, 2009
KIOLOARIA Harris V.A., 1994
Kioloaria bipunctata (Harris V.A., 2014)
Kioloaria brevicaudata (Thompson I.C. & Scott A., 1903)
Kioloaria cumula (Harris V.A., 2014)
Kioloaria edenensis (Harris V.A., 2014)
Kioloaria geddesi (Harris V.A., 2014)
Kioloaria iwasakii (Harris V.A., 2014)
Kioloaria jejuensis Huys, 2016
Kioloaria paguri (Ho, 1986)
Kioloaria quinquelineata (Harris V.A. & Robertson H.M., 1994)
Kioloaria rufolineata (Harris V.A. & Robertson, 1994)
Kioloaria sesquimaculata Harris V.A., 1994
Kioloaria similis (Kim S.H. & Kim W., 1996)
Kioloaria tapui (Hicks & Webber, 1983)
Key to species of Kioloaria Harris, 1994
KUSHIA Harris V.A. & Iwasaki, 1996
Kushia gamoi Harris V.A. & Iwasaki, 1996
Kushia igaguria Harris V.A. & Iwasaki, 1996
Kushia spathoides Harris V.A., 2014
Kushia zosteraphila Harris V.A. & Iwasaki, 1996 MUCROROSTRUM Harris V.A. & Iwasaki, 1997
Mucrorostrum yoroium Harris V.A. & Iwasaki, 1997
MURRAMIA Huys, 2016 (syn. Murramia Harris V.A., 1994)
Murramia bicincta Harris V.A., 1994
Murramia magna (Harris V.A., 1994)
Murramia poorei (Walker-Smith, 2001)
Subfamily Porcellidiinae Boeck, 1865
PORCELLIDIUM Claus, 1860 (syn. Thyone Philippi, 1840 )
Porcellidium acutum Kim I.H. & H.S. Kim, 1997 (syn.Kensakia acuta (Kim I.H. & Kim H.S., 1997)
Porcellidium acuticaudatum Thompson I.C. & Scott A., 1903 (syn. Kensakia acuticaudata (Thompson I.C. & Scott A., 1903)
Porcellidium affinis Quidor, 1906 (syn. Porcellidium affine Quidor, 1906
Porcellidium aiiroa (Harris V.A. & Iwasaki, 1997) (syn. Kensakia acuta (Kim I.H. & Kim H.S., 1997)
Porcellidium akashimum Harris V.A. & Iwasaki, 1996
Porcellidium algoense Hicks, 1982
Porcellidium aoifuchidorum Harris V.A. & Iwasaki, 1997 (syn. Porcellidium wandoensis Kim I.H. & H.S. Kim, 1997
Porcellidium bicincta (Harris V.A., 1994) (syn. Murramia bicincta Harris V.A., 1994
Porcellidium bipartitum Kim I.H. & H.S. Kim, 1997 (syn. Dilatatiocauda bipartita (Kim I.H. & H.S. Kim, 1997)
Porcellidium brevicaudatum Thompson I.C. & Scott A., 1903 (syn. Acutiramus brevicaudatus (Thompson I.C. & Scott A., 1903)
Porcellidium brevicavum Kim I.H. & H.S. Kim, 1997
Porcellidium clavigerum Pesta, 1935 (syn. Clavigofera clavigera (Pesta, 1935)
Porcellidium dentatum Claus, 1860 (syn. Porcelloides tenuicaudus (Claus, 1860)
Porcellidium dilatatum Hicks, 1971 (syn. Dilatatiocauda dilatata (Hicks, 1971)
Porcellidium echinophilum Humes & Gelerman, 1962 (syn. Clavigofera echinophila (Humes & Gelerman, 1962)
Porcellidium erythrogastrum Harris V.A. & H.M. Robertson, 1994
Porcellidium erythrum Hicks, 1971
Porcellidium fasciatum Boeck, 1865
Porcellidium fimbriatum Claus, 1863 (syn.Porcellidium fimbriatum fimbriatum Claus, 1863)
Porcellidium fulvum Thomson G.M., 1883 (syn.Porcellidium clavigerum Pesta, 1935)
Porcellidium gamoi (Harris V.A. & Iwasaki, 1996) (syn. Kushia gamoi Harris V.A. & Iwasaki, 1996
Porcellidium hartmannorum Tiemann, 1978
Porcellidium hormosirii Harris V.A. & H.M. Robertson, 1994
Porcellidium kiiroum Harris V.A. & Iwasaki, 1996
Porcellidium laurencium Hicks, 1982 (syn. Clavigofera laurencia (Hicks, 1982)
Porcellidium lecanoides Claus, 1889 (syn. Porcellidium viride (Philippi, 1840)
Porcellidium londonarum Harris V.A., 1994
Porcellidium londonii Harris, 1994 (?)
Porcellidium magna Harris V.A., 1994 (syn. Murramia magna (Harris V.A., 1994)
Porcellidium naviculum Harris V.A. & H.M. Robertson, 1994
Porcellidium ocellum Harris V.A. & Robertson H.M., 1994
Porcellidium ofunatense Harris V.A. & Iwasaki, 1996
Porcellidium ovatum Haller, 1879 (syn. Porcellidium tenuicauda Claus, 1860)
Porcellidium paguri Ho, 1986 (syn. Kioloaria paguri (Ho, 1986)
Porcellidium peniculiferum Tiemann, 1978 (syn. Porcellidium tristanense Wiborg, 1964)
Porcellidium phyllosporum Harris V.A. & H.M. Robertson, 1994
Porcellidium planum Tiemann, 1977 (syn. Dilatatiocauda plana (Tiemann, 1977)
Porcellidium poorei Walker-Smith, 2001 (syn. Murramia poorei (Walker-Smith, 2001)
Porcellidium pulchrum Harris V.A. & H.M. Robertson, 1994
Porcellidium quinquelineatus (Harris V.A. & Robertson H.M., 1994) (syn. Acutiramus quinquelineatus Harris V.A. & Robertson H.M., 1994)
Porcellidium rastellum Harris V.A., 2014
Porcellidium roscoffensis (Bocquet, 1948)
Porcellidium ravanae Thompson I.C. & Scott A., 1903 (syn.Ravania ravanae (Thompson I.C. & Scott A., 1903)
Porcellidium rufolineatus (Harris V.A. & Robertson H.M., 1994) (syn. Acutiramus rufolineatus Harris V.A. & Robertson H.M., 1994)
Porcellidium rubrum Pallares, 1966
Porcellidium sarsi Bocquet, 1948 (syn. Porcellidium viride (Philippi, 1840)
Porcellidium scutatum Claus, 1889 (syn. Porcelloides scutatus (Claus, 1889)
Porcellidium sesquimaculata Harris V.A., 1994 (syn. Acutiramus sesquimaculatus (Harris V.A., 1994)
Porcellidium similis Kim S.H. & Kim W., 1996 (syn. Acutiramus similis (Kim S.H. & Kim W., 1996)
Porcellidium tapui Hicks & Webber, 1983 (syn. Acutiramus tapui (Hicks & Webber, 1983)
Porcellidium tenuicauda Claus, 1860 (syn. Porcelloides tenuicaudus (Claus, 1860)
Porcellidium trisetosum Geddes, 1968 (syn. Geddesia trisetosa (Geddes, 1968)
Porcellidium tristanense Wiborg, 1964 (syn. Dilatatiocauda tristanensis (Wiborg, 1964)
Porcellidium ulvum Hicks, 1982 (syn. Clavigofera ulva (Hicks, 1982)
Porcellidium unicus Ummerkutty, 1970 (syn. Synurus unicus (Ummerkutty, 1970)
Porcellidium viridis (Philippi, 1840) (syn. Porcellidium viride (Philippi, 1840)
Porcellidium yoroium (Harris V.A. & Iwasaki, 1997) (syn. Mucrorostrum yoroium Harris V.A. & Iwasaki, 1997)
Porcellidium wandoensis Kim I.H. & H.S. Kim, 1997
PORCELLOIDES Harris V.A., 2014
Porcelloides scutatus (Claus, 1889)
Porcelloides tenuicaudus (Claus, 1860)
RAVANIA Harrris V.A., 2014
Ravania doliocauda Harris V.A., 2014
Ravania ravanae (Thompson I.C. & Scott A., 1903)
Ravania wellsi Harris V.A., 2014
SYNURUS Harris V.A., 2014
Synurus ctenocheirus Harris V.A., 2014
Synurus unicus (Ummerkutty, 1970)
TECTACIGULUM Huys, 2009
Tectacingulum nigrum (Harris V.A., 1994)
Tectacingulum tumidum (Harris V.A., 1994)
Acutiramus Harris V.A. & H.M. Robertson, 1994 (= Kioloaria Harris V.A., 1994 (Junior homonym);
Murramia Harris V.A., 1994 (= Murramia Huys, 2016;
Thyone Philippi, 1840 (=Porcellidium Claus, 1860 (synonym)
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