SUBFAMILY PARAMESOCHRINAE Huys, 1987


REMANEA Klie, 1929

  • Remanea arenicola Klie, 1929 [Germany; marine sand]
  • Remanea naksanensis Back et al., 2011 [Korea; brackish waters]
    Back & Lee (20129 reported this species as P. naksana
  • Remanea plumosa Pennak, 1942


    PARECHINOLAOPHONTE Song et al. 2023

    SCOTTOPSYLLUS Kunz, 1962

    "Kunz (1962) revised the Paramesochridae and introduced the new genera Scottopsyllus Kunz, 1962 and Intermedopsyllus Kunz, 1962. Later, Kunz (1981) combined the two genera and presented Scottopsyllus with three subgenera: Scottopsyllus, Intermedopsyllus and the newly established Wellsopsyllus Kunz, 1981. The genus group name Scottopsyllus as proposed by Kunz (1962), was declared as unavailable (nomen nudum) by Huys (2009) because he found ‘‘no trace of a type fixation in the original publication’’. Huys (2009) therefore replaced the name Scottopsyllus by the next oldest available name from among its synonyms, Wellsopsyllus Kunz, 1981. He found the same problem of lacking type fixation for the subgenus Intermedopsyllus. In the course of his nomenclatorical revision, Huys (2009) renamed the subgenera Scottopsyllus Apostolov and Marinov, 1988 and Intermediopsyllus Huys, 2009 (and introduced the new spelling Intermediopsyllus instead of Intermedopsyllus). At present, the genus Wellsopsyllus contains 15 species. The subgenus Wellsopsyllus Kunz, 1981 was conserved and contains 4 species to date, including the deep-sea species Wellsopsyllus (W.) abyssalis (Becker, 1979) and Wellsopsyllus (W.) antarcticus" (from: Kottmann et al., 2012)

    "The genus Scottopsyllus has been the subject of several revisions (Kunz 1962, 1981) and assessments of its status within the family of Paramesochridae (Kunz 1981, Huys 1987). Its division into the three subgenera Scottopsyllus, Intermedopsyllus and Wellsopsyllus is based only on diagnostic characters and may not reflect a phylogenetic affinity. Many important characters are still not taken into account and surely a revision will have to include not only the genus Scottopsyllus, but the Paramesochridae as a whole" (from: Veit-Kohler, 2021)


  • Wellsopsyllus herdmani (Thompson & A. Scott, 1899 [Europe; marine sand]
  • Wellsopsyllus langi langi Mielke, 1984 [Galapagos; marine sand] (=Wellsopsyllus (Scottopsyllus) langi (Mielke, 1984)
  • Wellsopsyllus langi continentalis Kun, 1992 [Yugoslavia; marine sand]
  • Wellsopsyllus minor (T. & A. Scott, 1895) [Europe; marine sand]
  • Wellsopsyllus pararobertsoni (Lang, 1965) [USA; marine sand]
  • Wellsopsyllus robertsoni (T. & A. Scott, 1895) [Europe; marine sand]
  • Wellsopsyllus antoniae (Plum & Kai-Horse, 2009) [North East Atlantic]
  • Wellsopsyllus intermedius (T. & A. Scott, 1895) [north Europe, Africa; Amphioxus sand]
  • Wellsopsyllus minutus (Nicholls, 1939) [Balloch Bay; marine sand]
  • Wellsopsyllus smirnovi Kunz, 1992 [Yugoslavia; marine sand]
  • Wellsopsyllus (Intermediopsyllus) minutus (Nicholls, 1939)
  • Wellsopsyllus (Intermediopsyllus) intermedius (Scott, T. & Scott A., 1895)
  • Wellsopsyllus (Intermediopsyllus) smirnovi (Kunz, 1992
  • Scottopsyllus (Intermedopsyllus) Kunz 1962 sensu Kunz [Northeast Atlantic]
  • Wellsopsyllus (Scottopsyllus) koreanus Back & Lee, 2014 [Korea; Yellow Sea]
  • Wellsopsyllus (Scottopsyllus) depressus (Kornev & Chertoprud, 2008
  • Wellsopsyllus (Scottopsyllus) herdmani (Thompson & Scott, 1900
  • Wellsopsyllus (Scottopsyllus) minor (Scott, T. & Scott A., 1895)
  • Wellsopsyllus (Scottopsyllus) pararobertsoni (Lang, 1965)
  • Wellsopsyllus (Scottopsyllus) praecipuus (Veit-Köhler, 2021)
  • Wellsopsyllus (Scottopsyllus) langi langi (Mielke, 1984)
  • Wellsopsyllus (Scottopsyllus) langi continentalis Kunz, 1992 [Yugoslavia]
  • Scottopsyllus runtzi (Soyer, 1975) accepted as Wellsopsyllus (Wellsopsyllus) runtzi (Soyer, 1975)
  • Wellsopsyllus (Scottopsyllus) robertsoni (Scott, T. & Scott A., 1895
  • Wellsopsyllus (Wellsopsyllus) abyssalis (Becker, Noodt & Schriever, 1979) [south America; marine sand]
  • Wellsopsyllus (Wellsopsyllus) gigas (Wells, 1965)
  • Wellsopsyllus (Wellsopsyllus) antarcticus Kottmann, Kihara, Glatzel & Veit-Koehler, 2013
  • Wellsopsyllus egregius Back & Lee, 2014 [Korea;Yellow Sea]
  • Wellsopsyllus koreanus Back & W. Lee, 2014
  • Wellsopsyllus (Wellsopsyllus) gigas (Wells, 1965) [Europe; marine sand]
  • Wellsopsyllus (Wellsopsyllus) runtzi (Soyer, 1975) [Kerguelen; mesopsammon]
  • Wellsopsyllus (Wellsopsyllus) antarcticus Kottmann et al., 2012 [Indian Ocean]


    SINGULARIA Huys, 1995 (SYN. OF BIUNCUS Huys, 1996)

  • Singularia ingens Huys, 1995 [Israel; Amphioxus sand]


    SUBFAMILY DIARTHRODELLINAE Huys, 1987


    TISBISOMA Bozic, 1964

    The genus Tisbisoma occupies an uncertain position among harpacticoids (Bozic, 1964). Kunz (19819 and Huys (1987) allocated it to the family Paramesochridae and, together with the genus Diarthrodella, to the subfamily Diarthrodellinae.

  • Tisbisoma spinisetum Bozic, 1964 [Reunion Island; marine]
  • Tisbisoma triarticulatum Wells, 1967 [Africa, marine]


    DIARTHRODELLA Klie, 1949

  • Diarthrodella chilensis Mielke, 1985 [Chile; marine]
  • Diarthrodella convexa Kunz, 1981 [Azores; marine]
  • Diarthrodella ergeneae Sonmez et al., 2014 [Turkey; sea waters]
  • Diarthrodella galapagoensis [Galapagos; marine interstitial]
  • Diarthrodella lancifera Kunz, 1981 [Azores; marine]
  • Diarthrodella neotropica Mielke, 1984 [Galapagos; marine interstitial]
  • Diarthrodella orbiculata Klie, 1949 [Helgoland; marine]
  • Diarthrodella parorbiculata parorbiculata Welsl, 1963 [north Ireland; marine]
  • Diarthrodella parorbiculata pacifica [Galapagos; marine interstitial]
  • Diarthrodella psammophila (Bocquet & Bozic, 1955) [France; marine sand]
  • Diarthrodella secunda secunda Kunz, 1954 [Germany; marine]
  • Diarthrodella secunda pacifica Kunz, 1981 [Pacific Ocean]

    |A key to the species of Diarthrodella |


    ROSSOPSYLLUS Soyer, 1975

  • Rossopsyllus kerguelensis kerguelensis Soyer, 1975 [Kerguelen Archipelago; marine]
  • Rossopsyllus kerguelensis quellonensis Mielke, 1985 [Chile; marine]
  • Rossopsyllus obscurus Cottarelli & Baldari, 1987 [Macquarie Island; marine interstitial]




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