BATHYPONTIDAE Brodsky, 1950


ALLOIOPODUS Bradford, 1969

  • Alloiopodus pinguis Bradford, 1969

    TEMORITES Sars G.O., 1900

    (syn. Bathypontia Sars G.O., 1905; Isocalanus Wolfenden, 1906 )

  • Temorites brevis Sars G.O., 1900
  • Temorites discoveryae Grice & Hulsemann, 1965
  • Temorites elegans (Sars G.O., 1920)
  • Temorites elongata (Sars G.O., 1905)
  • Temorites intermedia (Deevey, 1973)
  • Temorites kanaevae (Björnberg T.K.S., 1976)
  • Temorites longicornis (Tanaka, 1965)
  • Temorites longiseta (Brodsky, 1950)
  • Temorites michelae (Deevey, 1979)
  • Temorites minor (Wolfenden, 1906)
  • Temorites regalis (Grice & Hulsemann, 1967)
  • Temorites sarsi (Grice & Hulsemann, 1967)
  • Temorites similis (Tanaka, 1965)
  • Temorites spinifera (Scott A., 1909)
  • Temorites unispina (Deevey, 1979)

    TEMOROPSIS Wolfenden, 1911

  • Temoropsis simplex Wolfenden, 1911

    ZENKEVITCHIELLA Brodsky, 1955

  • Zenkevitchiella abyssalis Brodsky, 1955
  • Zenkevitchiella atlantica Grice & Hulsemann, 1965
  • Zenkevitchiella crassa Grice & Hulsemann, 1967
  • Zenkevitchiella tridentae Wheeler, 1970



    THARYBIDAE Sars G.O., 1902

    NEOSCOLECITHRIX Canu, 1896 (syn. Oothrix Farran, 1905 )

  • Neoscolecithrix antarctica Hulsemann, 1985 (syn. Cenognatha antarctica (Hulsemann, 1985)
  • Neoscolecithrix bidentata (Farran, 1905) (syn. Neoscolecithrix koehleri Canu, 1896)
  • Neoscolecithrix borealis (Wiborg, 1949)
  • Neoscolecithrix caetanoi Alvarez, 1985
  • Neoscolecithrix farrani Smirnov, 1935
  • Neoscolecithrix caetanoi Jimenez Alvares, 1985 [Brazil] (?)
  • Neoscolecithrix japonica Ohtsuka, Boxshall & Fosshagen, 2003
  • Neoscolecithrix koehleri Canu, 1896
  • Neoscolecithrix magna (Grice, 1973)
  • Neoscolecithrix ornata Bradford-Grieve, 2001
  • Neoscolecithrix watersae (Grice, 1973)

    PARUNDINELLA Fleminger, 1957

  • Parundinella dakini Bradford, 1973
  • Parundinella emarginata Grice & Hulsemann, 1970
  • Parundinella manicula Fleminger, 1957
  • Parundinella spinodenticula Fleminger, 1957

    RYTHABIS Schulz & Beckmann, 1995

  • Rythabis asymmetrica Markhaseva & Schulz, 2007
  • Rythabis atlantica Schulz & Beckmann, 1995
  • Rythabis heptneri Markhaseva & Ferrari, 2005
  • Rythabis schulzi Markhaseva & Ferrari, 2005

    THARIBIS Sars G.O., 1902 (syn. Tharybis compacta (Grice & Hulsemann, 1970)

    "Tharybis Sars, 1902 was established to accommodate Tharybis macrophthalma Sars, 1902 which was collected at about 180 m depth in the Christiania Fjord in Norway (Ferrari & Markhaseva, 2005). Andronov (2002) mentioned that species of the genus Tharybis mostly live near the bottom in depths from 30 to 4745 m and are rarely found in plankton samples. Since Sars (1902), in total 21 calanoid species were described belonging to Tharybis worldwide"(from Wang & Hwang, 2022 )

  • Tharybis altera (Grice & Hulsemann, 1970)
  • Tharybis angularis Schulz, 1995
  • Tharybis asymmetrica Andronov, 1976
  • Tharybis compacta (Grice & Hulsemann, 1970) (syn. Tharybis compacta (Grice & Hulsemann, 1970)
  • Tharybis crenata Schulz, 1995 (syn.Tharybis groenlandica (Tupitzky, 1982)
  • Tharybis fultoni Park, 1967
  • Tharybis groenlandica (Tupitzky, 1982)
  • Tharybis inaequalis Bradford-Grieve, 2001
  • Tharybis inflata Andronov, 2003
  • Tharybis juhlae Ferrari F.D. & Markhaseva, 2005
  • Tharybis kueishanensis Wang & Hwang, 2022 [Taiwan]
  • Tharybis lauta Andronov, 2003
  • Tharybis macrophthalamoida Andronov, 2003 (syn. Tharybis macrophthalmoida Andronov, 2003)
  • Tharybis macrophthalma Sars G.O., 1902
  • Tharybis macrophthalmoida Andronov, 2003
  • Tharybis magna Bradford & Wells, 1983
  • Tharybis megalodactyla Andronov, 1976
  • Tharybis minor Schulz, 1981 (syn. Tharybis asymmetrica Andronov, 1976)
  • Tharybis neptuni (Cleve, 1904)
  • Tharybis paraincertus (Grice & Hulsemann, 1965) (syn. Xanthocalanus paraincertus Grice & Hulsemann, 1965)
  • Tharybis pseudomegalodactyla Ferrari F.D. & Markhaseva, 2005
  • Tharybis sagamiensis Tanaka, 1960
  • Tharybis scaura Andronov, 2003
  • Tharybis shuheiella Ferrari F.D. & Markhaseva, 2005
  • Tharybis tuberosa Andronov, 2003
  • Tharybis tumidula Andronov, 2003


    UNDINELLA Sars G.O., 1900 (syn. Paratharybis Tanaka, 1937)

  • Undinella aculeata Andronov, 2003
  • Undinella acuta Vaupel Klein, 1970
  • Undinella altera Grice & Hulsemann, 1970 (syn. Tharybis altera (Grice & Hulsemann, 1970)
  • Undinella brevipes Farran, 1908 (syn. Undinella simplex (Wolfenden, 1906)
  • Undinella compacta Grice & Hulsemann, 1970
  • Undinella frontalis (Tanaka, 1937)
  • Undinella gricei Wheeler, 1970
  • Undinella hampsoni Grice & Hulsemann, 1970
  • Undinella hispidus (Grice & Hulsemann, 1967) (syn.Xanthocalanus hispidus Grice & Hulsemann, 1967)
  • Undinella messinensis (Claus, 1863) (syn. Euchirella messinensis (Claus, 1863)
  • Undinella oblonga Sars G.O., 1900
  • Undinella paululus (Park, 1970) (syn. Brodskius paululus (Park, 1970)
  • Undinella simplex (Wolfenden, 1906)
  • Undinella spinifer Tanaka, 1960
  • Undinella stirni Grice, 1971



    CLAUSOCALANIDAE Giesbrecht, 1893 (syn. Pseudocalanidae Giesbrecht, 1895)

    Clausia Boeck, 1865 accepted as Pseudocalanus Boeck, 1872; Drepanopsis Wolfenden, 1911 accepted as Farrania Sars G.O., 1920; Lucullus Giesbrecht, 1881 accepted as Pseudocalanus Boeck, 1872; Streptocalanus Brady, 1918 accepted as Pseudocalanus Boeck, 1872

    CLAUSOCALANUS Giesbrecht, 1888

  • Clausocalanus arcuicornis (Dana, 1849)(=Clausocalanus arcuicornis arcuicornis (Dana, 1849)
  • Clausocalanus brevipes Frost & Fleminger, 1968
  • Clausocalanus dubius Brodsky, 1950
  • Clausocalanus farrani Sewell, 1929
  • Clausocalanus furcatus (Brady, 1883)
  • Clausocalanus ingens Frost & Fleminger, 1968
  • Clausocalanus jobei Frost & Fleminger, 1968
  • Clausocalanus laticeps Farran, 1929
  • Clausocalanus lividus Frost & Fleminger, 1968
  • Clausocalanus mastigophorus (Claus, 1863)
  • Clausocalanus minor Sewell, 1929
  • Clausocalanus parapergens Frost & Fleminger, 1968
  • Clausocalanus paululus Farran, 1926
  • Clausocalanus pergens Farran, 1926

    CTENOCALANUS Giesbrecht, 1888

  • Ctenocalanus campaneri Almeida Prado-Por, 1984
  • Ctenocalanus citer Heron & Bowman, 1971
  • Ctenocalanus heronae Vega-Pérez & Bowman, 1992
  • Ctenocalanus longicornis Mori, 1937 (syn. Ctenocalanus vanus Giesbrecht, 1888
  • Ctenocalanus tageae Almeida Prado-Por, 1984
  • Ctenocalanus vanus Giesbrecht, 1888

    DREPANOPUS Brady, 1883

  • Drepanopus bispinosus Bayly, 1982
  • Drepanopus furcatus Brady, 1883 (=Clausocalanus furcatus (Brady, 1883)
  • Drepanopus bungei Sars G.O., 1898
  • Drepanopus forcipatus Giesbrecht, 1888
  • Drepanopus pectinatus Brady, 1883

    MICROCALANUS Sars G.O., 1903

  • Microcalanus cultrifer Farran, 1908
  • Microcalanus pusillus Sars G.O., 1903
  • Microcalanus pygmaeus (Sars G.O., 1900)

    PENICULOIDES Markhaseva & Renz, 2015

  • Peniculoides secundus Markhaseva & Renz, 2015

    PSEUDOCALANUS Boeck, 1872 (syn. Clausia Boeck, 1865; Lucullus Giesbrecht, 1881 Streptocalanus Brady, 1918)

  • Pseudocalanus acuspes (Giesbrecht, 1881)
  • Pseudocalanus armatus Boeck, 1872 (=Aetideus armatus (Boeck, 1872: Bradyidius armatus (Vanhöffen, 1897)
  • Pseudocalanus elongatus (Brady, 1865)
  • Pseudocalanus gracilis Sars G.O., 1903 (=Pseudocalanus minutus (Krøyer, 1845)
  • Pseudocalanus major Sars G.O., 1900 ( = Pseudocalanus minutus (Krøyer, 1845)
  • Pseudocalanus mimus Frost, 1989
  • Pseudocalanus minutus (Krøyer, 1845)
  • Pseudocalanus moultoni Frost, 1989
  • Pseudocalanus newmani Frost, 1989
  • Pseudocalanus pygmaeus Sars G.O., 1900 (=Microcalanus pygmaeus (Sars G.O., 1900)

    SPICIPES Grice & Hulsemann, 1965

  • Spicipes nanseni Grice & Hulsemann, 1965



    DIAPTOMIDAE Baird, 1850

    ACANTHODIAPTOMUS Kiefer, 1932

  • Acanthodiaptomus dampfi (Brehm, 1932) (syn. Arctodiaptomus dorsalis (Marsh, 1907)
  • Acanthodiaptomus denticornis (Wierzejski, 1887)
    Subspecies Acanthodiaptomus denticornis molvensis Monti & Stella, 1934
  • Acanthodiaptomus pacificus (Burckhardt, 1913)
    Subspecies Acanthodiaptomus pacificus yamanacensis
  • Acanthodiaptomus tibetanus (Daday, 1907)
  • Acanthodiaptomus yamanacensis (Brehm, 1925) (syn. Acanthodiaptomus pacificus (Burckhardt, 1913)

    AGLAODIAPTOMUS Light, 1938

  • Aglaodiaptomus atomicus DeBiase & Taylor, 1997
  • Aglaodiaptomus savagei DeBiase & Taylor, 2000
  • Aglaodiaptomus clavipes (Schacht, 1897)
  • Aglaodiaptomus clavipoides Wilson M.S., 1955
  • Aglaodiaptomus conipedatus (Marsh, 1907)
  • Aglaodiaptomus dilobatus Wilson M.S., 1958
  • Aglaodiaptomus forbesi Light, 1938
  • Aglaodiaptomus kingsburyae (Robertson A., 1975)
  • Aglaodiaptomus leptopus (Forbes S.A., 1882)
  • Aglaodiaptomus leptopus (Forbes S.A., 1882)
  • Aglaodiaptomus lintoni (Forbes S.A., 1893)
  • Aglaodiaptomus marshianus Wilson M.S., 1953
  • Aglaodiaptomus pseudosanguineus (Turner, 1921)
  • Aglaodiaptomus saskatchewanensis Wilson M.S., 1958
  • Aglaodiaptomus spatulocrenatus (Pearse, 1906)
  • Aglaodiaptomus savagei DeBiase & Taylor, 2000
  • Aglaodiaptomus stagnalis (Forbes S.A., 1882)

    ALLODIAPTOMUS Kiefer, 1936

    "The genus Allodiaptomus Kiefer, 1936 is a small group of freshwater diaptomid copepods that consists of five known species (Ranga Reddy et al., 2000; Walter and Boxshall, 2018), distributed mainly in river systems, reservoirs, and fish ponds (Ranga Reddy, 1987)"

  • Allodiaptomus calcarus Shen & Tai, 1965 (syn. Mongolodiaptomus calcarus (Shen & Tai, 1965)
  • Allodiaptomus cinctus (Gurney, 1907) (syn. Heliodiaptomus cinctus (Gurney, 1907)
  • Allodiaptomus gladiolus Shen & Lee, 1963 (syn. Mongolodiaptomus gladiolus (Shen & Lee, 1963)
  • Allodiaptomus intermedius Ranga Reddy, 1987
  • Allodiaptomus mieni Dang & Ho, 1985
  • Allodiaptomus mirabilipes Kiefer, 1936
  • Allodiaptomus nongensis Sanoamuang, L. & S. Watiroyram, 2019
  • Allodiaptomus pectinidactylus Shen & Tai, 1964 (syn. Mongolodiaptomus pectinidactylus (Shen & Tai, 1964)
  • Allodiaptomus raoi Kiefer, 1936
  • Allodiaptomus rappeportae (Brehm, 1923) (syn. Phyllodiaptomus rappeportae (Brehm, 1923)
  • Allodiaptomus rarus Ranga Reddy, Sanoamuang & Dumont, 1998
  • Allodiaptomus satanas (Brehm, 1952)
  • Allodiaptomus specillodactylus Shen & Tai, 1964 (syn. Allodiaptomus raoi Kiefer, 1936
  • Allodiaptomus tiruttani Rajendran, 1979 (possible syn. of Heliodiaptomus cinctus (Gurney, 1907)
  • Allodiaptomus uenoi (Kikuchi K., 1936) (syn. Mongolodiaptomus uenoi (Kikuchi K., 1936)

    ARCTODIAPTOMUS Kiefer, 1932

  • Arctodiaptomus (Pararctodiaptomus) Hsaio, 1950
  • Arctodiaptomus (Pararctodiaptomus) hsichowensis Hsiao, 1950
  • Arctodiaptomus (Rhabdodiapatomus) Kiefer, 1932
  • Arctodiaptomus (Stenodiaptomus) Kiefer, 1932
  • Arctodiaptomus acutilobatus acutilobatus (Sars G.O., 1903)
  • Arctodiaptomus acutilobatus curdicus Brehm, 1938
  • Arctodiaptomus acutulus (Brian, 1927)
  • Arctodiaptomus alpinus (Imhof, 1885)
  • Arctodiaptomus altissimus Kiefer, 1936
  • Arctodiaptomus altissimus pectinatus Shen & Sung, 1965
  • Arctodiaptomus anudarini Borutzky, 1959
  • Arctodiaptomus arapahoensis (Dodds, 1915)
  • Arctodiaptomus asymmetricus (Marsh, 1907)
  • Arctodiaptomus bacillifer (Koelbel, 1885)
  • Arctodiaptomus bacillifer proprior Kiefer, 1952
  • Arctodiaptomus bacillifer alpinus (Stella, 1984)
  • Arctodiaptomus bacillifer lagoraensis (Baldi,
  • Arctodiaptomus bacillifer montanus (Wierzejski,
  • Arctodiaptomus belgrati Mann, 1940
  • Arctodiaptomus brevirostris Dussart, 1974
  • Arctodiaptomus burduricus Kiefer, 1939
  • Arctodiaptomus byzantinus Mann, 1940
  • Arctodiaptomus centetes Brehm, 1938
  • Arctodiaptomus curdicus Brehm, 1938
  • Arctodiaptomus dahuricus Borutzky, 1959
  • Arctodiaptomus dentifer (Smirnov, 1928)
  • Arctodiaptomus dentifer rugosa Brehm, 1938
  • Arctodiaptomus dorsalis (Marsh, 1907)
  • Arctodiaptomus dudichi Kiefer, 1932
  • Arctodiaptomus euacanthus Kiefer, 1935
  • Arctodiaptomus faddeevi (Rylov, 1925) (syn. Arctodiaptomus spinosus (Daday, 1891)
  • Arctodiaptomus fischeri (Rylov, 1918)
  • Arctodiaptomus floridanus (Marsh, 1926)
  • Arctodiaptomus johanseni (Marsh, 1920) (syn. Limnocalanus johanseni Marsh, 1920
  • Arctodiaptomus jurisowitchi Löffler, 1968
  • Arctodiaptomus kamtschaticus Borutzky, 1953
  • Arctodiaptomus karadaghicus Ulomsky, 1941 (syn. Arctodiaptomus byzantinus Mann, 1940
  • Arctodiaptomus kerkyrensis Pesta, 1935
  • Arctodiaptomus kieferi Reddiah, 1965 (syn. Heliodiaptomus cinctus (Gurney, 1907)
  • Arctodiaptomus klebanovskyi Stepanova, 1999
  • Arctodiaptomus kliei Kiefer, 1933
  • Arctodiaptomus kupelwieseri (Brehm, 1907) (syn. Mixodiaptomus kupelwieseri (Brehm, 1907)
  • Arctodiaptomus kurilensis Kiefer, 1937
  • Arctodiaptomus laticeps (Sars G.O., 1863)
  • Arctodiaptomus lindbergi Brehm, 1959
  • Arctodiaptomus lisichowensis Shen, 1956
  • Arctodiaptomus lobulifer (Rylov, 1927)
  • Arctodiaptomus michaeli Ranga Reddy, Balkhi & Yousuf, 1990
  • Arctodiaptomus mildredi Streletskaya, 1986
  • Arctodiaptomus mucronatus Rylov, 1928 (syn. Arctodiaptomus acutulus (Brian, 1927)
  • Arctodiaptomus natronophilus Pónyi, 1956 (syn. Arctodiaptomus bacillifer (Koelbel, 1885)
  • Arctodiaptomus naurzumensis Stepanova, 1994
  • Arctodiaptomus nepalensis Uéno, 1966
  • Arctodiaptomus niethammeri niethammeri (Mann, 1940)
  • Arctodiaptomus niethammeri akatovae Kiefer, 1971
  • Arctodiaptomus novosibiricus Kiefer, 1971
  • Arctodiaptomus omskensis Stepanova, 1999
  • Arctodiaptomus osmanus Kiefer, 1974
  • Arctodiaptomus parvispineus Kiefer, 1935
  • Arctodiaptomus paulseni (Sars G.O., 1903)
  • Arctodiaptomus pectinicornis (Wierzejski, 1887)
  • Arctodiaptomus ohridanus Brehm, 1939 (syn. Arctodiaptomus stephanidesi stephanidesi (Pesta, 1935)
  • Arctodiaptomus piliger Brehm, 1955 (syn. Arctodiaptomus alpinus (Imhof, 1885)
  • Arctodiaptomus proximus (Kiefer, 1936) (syn. Arctodiaptomus dorsalis (Marsh, 1907)
  • Arctodiaptomus rectispinosus rectispinosus Kikuchi K., 1940
  • Arctodiaptomus rectispinosus mongolicus Flössner, 1986
  • Arctodiaptomus salinus (Daday, 1885)
  • Arctodiaptomus saltillinus (Brewer, 1898)
  • Arctodiaptomus sensibilis Mann, 1940 (syn. Arctodiaptomus stephanidesi
  • Arctodiaptomus shillongensis Reddiah, 1965 (syn. Heliodiaptomus cinctus (Gurney, 1907)
  • Arctodiaptomus similis (Baird, 1859)
  • Arctodiaptomus similis irregularis Kiefer, 1974
  • Arctodiaptomus similis saetosior Kiefer, 1978
  • Arctodiaptomus spectabilis Mann, 1940
  • Arctodiaptomus spinosus (Daday, 1891)
  • Arctodiaptomus spirulus Shen, 1956
  • Arctodiaptomus stephanidesi stephanidesi (Pesta, 1935)
  • Arctodiaptomus stephanidesi bulgaricus Kiefer, 1971
  • Arctodiaptomus steindachneri (Richard, 1897)
  • Arctodiaptomus steindachneri kerkyrensis Pesta, 1935
  • Arctodiaptomus stewartianus (Brehm, 1925)
  • Arctodiaptomus tianchiensis Chen & Hu, 1989
  • Arctodiaptomus tianchiensis Chen & Hu, 1989
  • Arctodiaptomus (Mesodiaptomus) toni Brehm, 1937
  • Arctodiaptomus ulomskyi Chechuro, 1960
  • Arctodiaptomus valachicus Banarescu & Serban, 1954 (syn. Arctodiaptomus dudichi Kiefer, 1932
  • Arctodiaptomus walterianus (Brehm, 1925)
  • Arctodiaptomus wierzejskii (Richard, 1888)
  • Arctodiaptomus winbergi Stepanova, 2001

    ARGYRODIAPTOMUS Brehm, 1933

  • Argyrodiaptomus aculeatus (Douwe, 1911)
  • Argyrodiaptomus argentinus (Wright S., 1938)
  • Argyrodiaptomus azevedoi (Wright S., 1935)
  • Argyrodiaptomus bergi (Richard, 1897)
  • Argyrodiaptomus denticulatus (Pesta, 1927)
  • Argyrodiaptomus exilis Dussart, 1985
  • Argyrodiaptomus falcifer (Daday, 1905) (syn. Diaptomus falcifer Daday, 1905)
  • Argyrodiaptomus ferus Shen & Tai, 1964 (syn. Sinodiaptomus ferus (Shen & Tai, 1964)
  • Argyrodiaptomus furcatus (Sars G.O., 1901)
  • Argyrodiaptomus furcatus exilis Dussart, 1985
  • Argyrodiaptomus furcatus meridionalis Dussart, 1985
  • Argyrodiaptomus furcatus septentrionalis Dussart, 1985
  • Argyrodiaptomus granulosus Brehm, 1933
  • Argyrodiaptomus macrochaetus Brehm, 1937
  • Argyrodiaptomus neglectus (Wright S., 1937)
  • Argyrodiaptomus nhumirim Reid, 1997
  • Argyrodiaptomus paggi Previattelli & Santos-Silva, 2007
  • Argyrodiaptomus robertsonae Dussart, 1985
  • Argyrodiaptomus spiniger (Brian, 1926)
  • Argyrodiaptomus cavernicolax Shen & Tai, 1965

    ASPINUS Brandorff, 1973

  • Aspinus acicularis Brandorff, 1973

    AUSTRINODIAPTOMUS Reid, 1997

  • Austrinodiaptomus kleerekoperi Reid, 1997

    CALODIAPTOMUS Kiefer, 1936

  • Calodiaptomus merrillae (Wright S., 1927)
  • Calodiaptomus perelegans (Wright S., 1927)

    CAMERUNDIAPTOMUS Dumont & Chiambeng, 2002

  • Camerundiaptomus christineae Dumont & Chiambeng, 2002
  • Camerundiaptomus djamai Dumont & Chiambeng, 2002

    COLOMBODIAPTOMUS Gaviria, 1989

  • Colombodiaptomus brandorffi Gaviria, 1989
  • Colombodiaptomus brandorffi pilosus Gaviria, 1989

    COPIDODIAPTOMUS Kiefer, 1968

  • Copidodiaptomus numidicus (Gurney, 1909)
  • Copidodiaptomus steueri (Brehm & Zederbauer, 1904)

    DACTYLODIAPTOMUS Kiefer, 1936

  • Dactylodiaptomus pearsei (Wright S., 1927)

    DASYDIAPTOMUS Defaye & Dussart, 1993 (syn. Trichodiaptomus Brandorff, Koste & Smirnov, 1982)

  • Dasydiaptomus coronatus (Sars G.O., 1901)

    DENTODIAPTOMUS Shen & Tai, 1964

  • Dentodiaptomus javanus (Grochmalicki, 1915)
  • Dentodiaptomus orientalis Sanoamuang & Watiroyram, 2021


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