MEGACALANIDAE Sewell, 1947
BATHYCALANUS Sars G.O., 1905
Bathycalanus adornatus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Bathycalanus bradyi (Wolfenden, 1905)
Bathycalanus bucklinae Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Bathycalanus dentatus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Bathycalanus eltaninae Björnberg T.K.S., 1968 (syn. Elenacalanus eltaninae (Björnberg, 1968)
Bathycalanus eximius Markhaseva, in Brodsky, Vyshkvartseva, Kos & Markhaseva, 1983
Bathycalanus inflatus Björnberg T.K.S., 1968 (syn. Elenacalanus inflatus (Björnberg, 1968)
Bathycalanus maximus Wolfenden, 1911 (syn. Bathycalanus bradyi (Wolfenden, 1905)
Bathycalanus milleri Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Bathycalanus princeps (Brady, 1883) (syn. Elenacalanus princeps (Brady, 1883)
Bathycalanus pustulosus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Bathycalanus richardi Sars G.O., 1905
Bathycalanus rigidus Sars G.O., 1920 (syn. Elenacalanus princeps (Brady, 1883)
Bathycalanus sverdrupi Johnson M.W., 1958 (syn. Elenacalanus sverdrupi (Johnson, 1958)
Bathycalanus tumidus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Bathycalanus unicornis Björnberg T.K.S., 1968
BRADYCALANUS Scott A., 1909
Bradycalanus abyssicolus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Bradycalanus enormis Björnberg, 1968
Bradycalanus gigas Sewell, 1947
Bradycalanus pseudotypicus Björnberg T.K.S., 1968 (syn. Bradycalanus typicus Scott A., 1909
Bradycalanus sarsi (Farran, 1939)(syn. Megacalanus sarsi Farran, 1939; Bradycalanus typicus Scott A., 1909
Bradycalanus typicus Scott A., 1909
ELENACALANUS Bradford-Grieve, Blanco-Bercial & Boxshall, 2017 (syn. Heterocalanus sensu Wolfenden, 1906)
Elenacalanus eltaninae (Björnberg, 1968)
Elenacalanus inflatus (Björnberg, 1968)
Elenacalanus princeps (Brady, 1883)
Elenacalanus sverdrupi (Johnson, 1958)
Elenacalanus tageae Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
MEGACALANUS Wolfenden, 1904 (syn. Macrocalanus Sars G.O., 1905)
Megacalanus bradyi Wolfenden, 1905 (syn. Bathycalanus bradyi (Wolfenden, 1905)
Megacalanus ericae Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Megacalanus frosti Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Megacalanus gracilis (Dana, 1852) (syn. Neocalanus gracilis (Dana, 1852)
Megacalanus longicornis (Sars G.O., 1905) (syn. Megacalanus princeps Wolfenden, 1904; Megacalanus princeps princeps Wolfenden, 1904)
Megacalanus ohmani Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
Megacalanus princeps Wolfenden, 1904 (syn. Megacalanus princeps princeps Wolfenden, 1904)
Megacalanus robustior (Giesbrecht, 1888) (syn. Neocalanus robustior (Giesbrecht, 1888)
Megacalanus sarsi Farran, 1939 (syn. Bradycalanus typicus Scott A., 1909)
Megacalanus typicus (Scott A., 1909)(syn.Bradycalanus typicus Scott A., 1909)
PARAPONTELLIDAE Giesbrecht, 1893
NEOPONTELLA Scott A., 1909
Neopontella typica Scott A., 1909
PARAPONTELLA Brady, 1878
Parapontella brevicornis (Lubbock, 1857)
PHAENNIDAE Sars G.O., 1902
BRACHYCALANUS Farran, 1905
"Brachycalanus is a genus of peculiar benthopelagic
copepods which bear close resemblance to
Xanthocalanus species. The genus was created by
Farran (1905) to include X. at/anticus, being
distinguished from true species of Xanthocalanus in
the shape of the rostrum and also the short first
antenna" (from:Othman & Greenwood. 1988)
Brachycalanus antarcticus Schulz, 2005
Brachycalanus atlanticus (Wolfenden, 1904)
Brachycalanus bjornbergae Campaner, 1978
Brachycalanus brodskyi Ferrari F.D. & Markhaseva, 2000
Brachycalanus flemingeri Ferrari F.D. & Markhaseva, 2000
Brachycalanus gigas Scott A., 1909 (syn. Landrumius gigas (Scott A., 1909)
Brachycalanus minutus Grice, 1973
Brachycalanus ordinarius (Grice, 1973)
Brachycalanus rothlisbergi Othman & Greenwood, 1988
CEPHALOPHANES Sars G.O., 1907
Cephalophanes frigidus Wolfenden, 1911
Cephalophanes refulgens Sars G.O., 1907
Cephalophanes tectus (Esterly, 1911)
CORNUCALANUS Wolfenden, 1905
Cornucalanus antarcticus Brodsky & Zvereva, 1976 (syn. Cornucalanus robustus Vervoort, 1957)
Cornucalanus chelifer (Thompson I.C., 1903)
Cornucalanus indicus Sewell, 1929
Cornucalanus magnus Wolfenden, 1905 (syn. Cornucalanus chelifer (Thompson I.C., 1903)
Cornucalanus notabilis Brodsky & Zvereva, 1976
Cornucalanus robustus Vervoort, 1957
Cornucalanus sewelli Vervoort, 1957
Cornucalanus simplex Wolfenden, 1905
KIRNESIUS Markhaseva & Semenova, 2005
Kirnesius groenlandicus Markhaseva & Semenova, 2005 ONCHOCALANUS Sars G.O., 1905
Onchocalanus affinis With, 1915
Onchocalanus chelifer (Thompson I.C., 1903) (syn.Cornucalanus chelifer (Thompson I.C., 1903)
Onchocalanus cristatus (Wolfenden, 1904)
Onchocalanus cristogerens Markhaseva & Kosobokova, 1998
Onchocalanus frigidus Wolfenden, 1911 (syn. Onchocalanus magnus (Wolfenden, 1906)
Onchocalanus hirtipes Sars G.O., 1905
Onchocalanus latus Esterly, 1911
Onchocalanus magnus (Wolfenden, 1906)
Onchocalanus nudipes Wilson C.B, 1942 (syn. Phaenna spinifera Claus, 1863)
Onchocalanus paratrigoniceps Park, 1983
Onchocalanus scotti Vervoort, 1950
Onchocalanus steueri Pesta, 1920 (syn. Onchocalanus trigoniceps Sars G.O., 1905)
Onchocalanus subcristatus Wolfenden, 1911
Onchocalanus trigoniceps Sars G.O., 1905
Onchocalanus wolfendeni Vervoort, 1950
PHENNA Claus, 1863
Phaenna gibbosa Andronov, 2013
Phaenna latus Esterly, 1911
Phaenna spinifera Claus, 1863
Phaenna zetlandica Scott T., 1902
PHAENNOCALANUS Markhaseva, 2002
Phaennocalanus unispinosus Markhaseva, 2002
TALACALANUS Wolfenden, 1911
Talacalanus calaminus Wolfenden, 1911
Talacalanus greenii (Farran, 1905)
Talacalanus maximus (Brodsky, 1950)
AMALLOPHORA Scott T., 1894 (= Xanthocalanus Giesbrecht, 1893)
XANTHOCALANUS Giesbrecht, 1893 (syn. Amallophora Scott T., 1894) The Phaennidae genus Xanthocalanus is the largest
among the genera of the Bradfordian families.
As much as 67 species were described in
Xanthocalanus (Razouls, 1995), but subsequently
17 species were tentatively or definitely transferred
to Tharybidae or Scolecitrichidae (Bradford,
1973; Roe, 1975; Campaner, 1978; Bradford
et al., 1983; Ohtsuka et al., 1998; Markhaseva,
1998; Vyshkvartzeva, 2000) (from Vyshkvartzeva, 2002 (=Scolecitrichopsis Vyshkvartzeva 2000; Pseudoamallothrix Vyshkvartzeva 2000
Xanthocalanus agilis Giesbrecht, 1893
Xanthocalanus alterus (Farran, 1929) (syn. Mixtocalanus alter (Farran, 1929)
Xanthocalanus alvinae Grice & Hulsemann, 1970
Xanthocalanus amabilis Tanaka, 1960
Xanthocalanus antarcticus Wolfenden, 1908
Xanthocalanus atlanticus Wolfenden, 1904 (syn.Brachycalanus atlanticus (Wolfenden, 1904)
Xanthocalanus borealis Sars G.O., 1900 (syn. Xanthocalanus hirtipes Vanhöffen, 1897
Xanthocalanus brevicornis (Sars G.O., 1900) (= Scaphocalanus brevicornis (Sars G.O., 1900)
Xanthocalanus calaminus Wolfenden, 1911 (= Talacalanus greenii (Farran, 1905)
Xanthocalanus claviger (Scott T., 1909)
Xanthocalanus cornifer (Tanaka, 1960)
Xanthocalanus crassirostris (Tanaka, 1960)
Xanthocalanus cryeri (Bradford-Grieve, 2005) (=Xantharus cryeri Bradford-Grieve, 2005)
Xanthocalanus cristatus Wolfenden, 1904 (=Onchocalanus cristatus (Wolfenden, 1904)
Xanthocalanus difficilis Grice & Hulsemann, 1965 (= Scolecitrichopsis difficilis (Grice & Hulsemann, 1965)
Xanthocalanus dilatus Grice, 1962
Xanthocalanus distinctus Grice & Hulsemann, 1970 (syn. Scolecitrichopsis distinctus (Grice & Hulsemann, 1970)
Xanthocalanus echinatus Sars G.O., 1907
Xanthocalanus elegans (Wolfenden, 1911) (= Scaphocalanus elegans (Wolfenden, 1911)
Xanthocalanus elongatus Grice & Hulsemann, 1970 (= Scolecitrichopsis elongatus (Grice & Hulsemann, 1970)
Xanthocalanus fallax Sars G.O., 1903
Xanthocalanus fragilis Aurivillius, 1898 (syn. Xanthocalanus minor Giesbrecht, 1893)
Xanthocalanus giesbrechti Thompson I.C., 1903
Xanthocalanus globiceps (Farran, 1908) (syn. Amallophora globiceps (Farran, 1908)
Xanthocalanus gracilis Wolfenden, 1911
Xanthocalanus greeni Farran, 1905 (syn. Xanthocalanus greenii Farran, 1905; Talacalanus greenii (Farran, 1905)
Xanthocalanus groenlandicus Tupitzky, 1982 (syn.Tharybis groenlandicus (Tupitzky, 1982): Tharybis groenlandica (Tupitzky, 1982)
Xanthocalanus harpagatus Bradford & Wells, 1983
Xanthocalanus hirtipes Vanhöffen, 1897
Xanthocalanus hispidus Grice & Hulsemann, 1967
Xanthocalanus impar (Wolfenden, 1911) (syn. Scaphocalanus impar (Wolfenden, 1911)
Xanthocalanus incertus Sars G.O., 1920
Xanthocalanus irritans (Tanaka, 1960)
Xanthocalanus kurilensis Brodsky, 1950
Xanthocalanus laptevorum Markhaseva, 1998 (= Byrathis laptevorum (Markhaseva, 1998)
Xanthocalanus legatus Tanaka, 1960
Xanthocalanus longispinus Bradford-Grieve, 2004
Xanthocalanus macilenta (Grice & Hulsemann, 1970)
Xanthocalanus macrocephalon Grice & Hulsemann, 1970 (syn.Tharybis macrocephalon (Grice & Hulsemann, 1970); Byrathis macrocephalon (Grice & Hulsemann, 1970)
Xanthocalanus magnus Wolfenden, 1906 (= Onchocalanus magnus (Wolfenden, 1906)
Xanthocalanus marlyae Campaner, 1978
Xanthocalanus maximus Brodsky, 1950 (syn.Talacalanus maximus (Brodsky, 1950)
Xanthocalanus medius Tanaka, 1937
Xanthocalanus meteorae Markhaseva & Schnack-Schiel, 2003
Xanthocalanus minor Giesbrecht, 1893
Xanthocalanus mixtus Sars G.O., 1920
Xanthocalanus multispinus Chen & Zhang, 1975
Xanthocalanus muticus Sars G.O., 1905
Xanthocalanus oculatus (Tanaka, 1960)
Xanthocalanus obtusus Farran, 1904
Xanthocalanus ordinarius Grice, 1973 (= Brachycalanus ordinarius (Grice, 1973)
Xanthocalanus paraincertus Grice & Hulsemann, 1965 (syn. Brodskius robustipes (Grice & Hulsemann, 1965)
Xanthocalanus paululus Park, 1970 (= Brodskius paululus (Park, 1970)
Xanthocalanus pavlovskii Brodsky, 1955
Xanthocalanus pectinatus Tanaka, 1960
Xanthocalanus penicillatus Tanaka, 1960
Xanthocalanus pinguis Farran, 1905
Xanthocalanus polaris Brodsky, 1950 (=Scolecitrichopsis polaris (Brodsky, 1950)
Xanthocalanus polarsternae Markhaseva, 1998
Xanthocalanus profundus Sars G.O., 1907
Xanthocalanus propinquus Sars G.O., 1902
Xanthocalanus ptevorum Markhaseva, 1998 [Laptev Sea]
Xanthocalanus pulcher Esterly, 1911
Xanthocalanus quasiprofundus Vyshkvartzeva, 2002
Xanthocalanus robustus (Scott T., 1894) (syn.Amallothrix robusta (Scott T., 1894)
Xanthocalanus rotundus (Grice & Hulsemann, 1970)
Xanthocalanus serratus (Tanaka, 1960)
Xanthocalanus similis Esterly, 1906 (syn. Onchocalanus cristatus (Wolfenden, 1904)
Xanthocalanus simplex Aurivillius, 1898 (syn.Undinella simplex (Wolfenden, 1906)
Xanthocalanus smithae (Grice, 1962) (syn. Scopalatum smithae (Grice, 1962)
Xanthocalanus soaresmoreirai Björnberg T.K.S., 1975 (syn. Pseudoamallothrix soaresmoreirai (Björnberg T.K.S., 1975)
Xanthocalanus spinodenticulatus Markhaseva, 1998 [Laptev Sea]
Xanthocalanus squamatus Farran, 1936
Xanthocalanus stewarti Bradford-Grieve, 2004
Xanthocalanus subagilis Wolfenden, 1904
Xanthocalanus subbrevicornis (syn. Scaphocalanus subbrevicornis (Wolfenden, 1911)
Xanthocalanus subcristatus Wolfenden, 1906 (syn. Onchocalanus subcristatus Wolfenden, 1911
Xanthocalanus tectus Esterly, 1911 (syn. Cephalophanes tectus (Esterly, 1911)
Xanthocalanus tenuiremis Scott T., 1909
Xanthocalanus tenuiserratus Wolfenden, 1911
Xanthocalanus typicus (Scott T., 1894)
Xanthocalanus validus (Farran, 1908) (syn.Amallothrix valida (Farran, 1908)
Xanthocalanus watersae Grice, 1973 (syn. Neoscolecithrix watersae (Grice, 1973)
Key to Megacalanidae genera
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