MEGACALANIDAE Sewell, 1947

BATHYCALANUS Sars G.O., 1905

  • Bathycalanus adornatus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Bathycalanus bradyi (Wolfenden, 1905)
  • Bathycalanus bucklinae Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Bathycalanus dentatus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Bathycalanus eltaninae Björnberg T.K.S., 1968 (syn. Elenacalanus eltaninae (Björnberg, 1968)
  • Bathycalanus eximius Markhaseva, in Brodsky, Vyshkvartseva, Kos & Markhaseva, 1983
  • Bathycalanus inflatus Björnberg T.K.S., 1968 (syn. Elenacalanus inflatus (Björnberg, 1968)
  • Bathycalanus maximus Wolfenden, 1911 (syn. Bathycalanus bradyi (Wolfenden, 1905)
  • Bathycalanus milleri Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Bathycalanus princeps (Brady, 1883) (syn. Elenacalanus princeps (Brady, 1883)
  • Bathycalanus pustulosus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Bathycalanus richardi Sars G.O., 1905
  • Bathycalanus rigidus Sars G.O., 1920 (syn. Elenacalanus princeps (Brady, 1883)
  • Bathycalanus sverdrupi Johnson M.W., 1958 (syn. Elenacalanus sverdrupi (Johnson, 1958)
  • Bathycalanus tumidus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Bathycalanus unicornis Björnberg T.K.S., 1968

    BRADYCALANUS Scott A., 1909

  • Bradycalanus abyssicolus Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Bradycalanus enormis Björnberg, 1968
  • Bradycalanus gigas Sewell, 1947
  • Bradycalanus pseudotypicus Björnberg T.K.S., 1968 (syn. Bradycalanus typicus Scott A., 1909
  • Bradycalanus sarsi (Farran, 1939)(syn. Megacalanus sarsi Farran, 1939; Bradycalanus typicus Scott A., 1909
  • Bradycalanus typicus Scott A., 1909

    ELENACALANUS Bradford-Grieve, Blanco-Bercial & Boxshall, 2017 (syn. Heterocalanus sensu Wolfenden, 1906)

  • Elenacalanus eltaninae (Björnberg, 1968)
  • Elenacalanus inflatus (Björnberg, 1968)
  • Elenacalanus princeps (Brady, 1883)
  • Elenacalanus sverdrupi (Johnson, 1958)
  • Elenacalanus tageae Bradford-Grieve, Blanco-Bercial & Boxshall, 2017

    MEGACALANUS Wolfenden, 1904 (syn. Macrocalanus Sars G.O., 1905)

  • Megacalanus bradyi Wolfenden, 1905 (syn. Bathycalanus bradyi (Wolfenden, 1905)
  • Megacalanus ericae Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Megacalanus frosti Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Megacalanus gracilis (Dana, 1852) (syn. Neocalanus gracilis (Dana, 1852)
  • Megacalanus longicornis (Sars G.O., 1905) (syn. Megacalanus princeps Wolfenden, 1904; Megacalanus princeps princeps Wolfenden, 1904)
  • Megacalanus ohmani Bradford-Grieve, Blanco-Bercial & Boxshall, 2017
  • Megacalanus princeps Wolfenden, 1904 (syn. Megacalanus princeps princeps Wolfenden, 1904)
  • Megacalanus robustior (Giesbrecht, 1888) (syn. Neocalanus robustior (Giesbrecht, 1888)
  • Megacalanus sarsi Farran, 1939 (syn. Bradycalanus typicus Scott A., 1909)
  • Megacalanus typicus (Scott A., 1909)(syn.Bradycalanus typicus Scott A., 1909)


    PARAPONTELLIDAE Giesbrecht, 1893

    NEOPONTELLA Scott A., 1909

  • Neopontella typica Scott A., 1909

    PARAPONTELLA Brady, 1878

  • Parapontella brevicornis (Lubbock, 1857)


    PHAENNIDAE Sars G.O., 1902

    BRACHYCALANUS Farran, 1905

    "Brachycalanus is a genus of peculiar benthopelagic copepods which bear close resemblance to Xanthocalanus species. The genus was created by Farran (1905) to include X. at/anticus, being distinguished from true species of Xanthocalanus in the shape of the rostrum and also the short first antenna" (from:Othman & Greenwood. 1988)

  • Brachycalanus antarcticus Schulz, 2005
  • Brachycalanus atlanticus (Wolfenden, 1904)
  • Brachycalanus bjornbergae Campaner, 1978
  • Brachycalanus brodskyi Ferrari F.D. & Markhaseva, 2000
  • Brachycalanus flemingeri Ferrari F.D. & Markhaseva, 2000
  • Brachycalanus gigas Scott A., 1909 (syn. Landrumius gigas (Scott A., 1909)
  • Brachycalanus minutus Grice, 1973
  • Brachycalanus ordinarius (Grice, 1973)
  • Brachycalanus rothlisbergi Othman & Greenwood, 1988

    CEPHALOPHANES Sars G.O., 1907

  • Cephalophanes frigidus Wolfenden, 1911
  • Cephalophanes refulgens Sars G.O., 1907
  • Cephalophanes tectus (Esterly, 1911)

    CORNUCALANUS Wolfenden, 1905

  • Cornucalanus antarcticus Brodsky & Zvereva, 1976 (syn. Cornucalanus robustus Vervoort, 1957)
  • Cornucalanus chelifer (Thompson I.C., 1903)
  • Cornucalanus indicus Sewell, 1929
  • Cornucalanus magnus Wolfenden, 1905 (syn. Cornucalanus chelifer (Thompson I.C., 1903)
  • Cornucalanus notabilis Brodsky & Zvereva, 1976
  • Cornucalanus robustus Vervoort, 1957
  • Cornucalanus sewelli Vervoort, 1957
  • Cornucalanus simplex Wolfenden, 1905

    KIRNESIUS Markhaseva & Semenova, 2005

  • Kirnesius groenlandicus Markhaseva & Semenova, 2005

    ONCHOCALANUS Sars G.O., 1905

  • Onchocalanus affinis With, 1915
  • Onchocalanus chelifer (Thompson I.C., 1903) (syn.Cornucalanus chelifer (Thompson I.C., 1903)
  • Onchocalanus cristatus (Wolfenden, 1904)
  • Onchocalanus cristogerens Markhaseva & Kosobokova, 1998
  • Onchocalanus frigidus Wolfenden, 1911 (syn. Onchocalanus magnus (Wolfenden, 1906)
  • Onchocalanus hirtipes Sars G.O., 1905
  • Onchocalanus latus Esterly, 1911
  • Onchocalanus magnus (Wolfenden, 1906)
  • Onchocalanus nudipes Wilson C.B, 1942 (syn. Phaenna spinifera Claus, 1863)
  • Onchocalanus paratrigoniceps Park, 1983
  • Onchocalanus scotti Vervoort, 1950
  • Onchocalanus steueri Pesta, 1920 (syn. Onchocalanus trigoniceps Sars G.O., 1905)
  • Onchocalanus subcristatus Wolfenden, 1911
  • Onchocalanus trigoniceps Sars G.O., 1905
  • Onchocalanus wolfendeni Vervoort, 1950

    PHENNA Claus, 1863

  • Phaenna gibbosa Andronov, 2013
  • Phaenna latus Esterly, 1911
  • Phaenna spinifera Claus, 1863
  • Phaenna zetlandica Scott T., 1902

    PHAENNOCALANUS Markhaseva, 2002

  • Phaennocalanus unispinosus Markhaseva, 2002

    TALACALANUS Wolfenden, 1911

  • Talacalanus calaminus Wolfenden, 1911
  • Talacalanus greenii (Farran, 1905)
  • Talacalanus maximus (Brodsky, 1950)

    AMALLOPHORA Scott T., 1894 (= Xanthocalanus Giesbrecht, 1893)

    XANTHOCALANUS Giesbrecht, 1893 (syn. Amallophora Scott T., 1894)

    The Phaennidae genus Xanthocalanus is the largest among the genera of the Bradfordian families. As much as 67 species were described in Xanthocalanus (Razouls, 1995), but subsequently 17 species were tentatively or definitely transferred to Tharybidae or Scolecitrichidae (Bradford, 1973; Roe, 1975; Campaner, 1978; Bradford et al., 1983; Ohtsuka et al., 1998; Markhaseva, 1998; Vyshkvartzeva, 2000) (from Vyshkvartzeva, 2002 (=Scolecitrichopsis Vyshkvartzeva 2000; Pseudoamallothrix Vyshkvartzeva 2000

  • Xanthocalanus agilis Giesbrecht, 1893
  • Xanthocalanus alterus (Farran, 1929) (syn. Mixtocalanus alter (Farran, 1929)
  • Xanthocalanus alvinae Grice & Hulsemann, 1970
  • Xanthocalanus amabilis Tanaka, 1960
  • Xanthocalanus antarcticus Wolfenden, 1908
  • Xanthocalanus atlanticus Wolfenden, 1904 (syn.Brachycalanus atlanticus (Wolfenden, 1904)
  • Xanthocalanus borealis Sars G.O., 1900 (syn. Xanthocalanus hirtipes Vanhöffen, 1897
  • Xanthocalanus brevicornis (Sars G.O., 1900) (= Scaphocalanus brevicornis (Sars G.O., 1900)
  • Xanthocalanus calaminus Wolfenden, 1911 (= Talacalanus greenii (Farran, 1905)
  • Xanthocalanus claviger (Scott T., 1909)
  • Xanthocalanus cornifer (Tanaka, 1960)
  • Xanthocalanus crassirostris (Tanaka, 1960)
  • Xanthocalanus cryeri (Bradford-Grieve, 2005) (=Xantharus cryeri Bradford-Grieve, 2005)
  • Xanthocalanus cristatus Wolfenden, 1904 (=Onchocalanus cristatus (Wolfenden, 1904)
  • Xanthocalanus difficilis Grice & Hulsemann, 1965 (= Scolecitrichopsis difficilis (Grice & Hulsemann, 1965)
  • Xanthocalanus dilatus Grice, 1962
  • Xanthocalanus distinctus Grice & Hulsemann, 1970 (syn. Scolecitrichopsis distinctus (Grice & Hulsemann, 1970)
  • Xanthocalanus echinatus Sars G.O., 1907
  • Xanthocalanus elegans (Wolfenden, 1911) (= Scaphocalanus elegans (Wolfenden, 1911)
  • Xanthocalanus elongatus Grice & Hulsemann, 1970 (= Scolecitrichopsis elongatus (Grice & Hulsemann, 1970)
  • Xanthocalanus fallax Sars G.O., 1903
  • Xanthocalanus fragilis Aurivillius, 1898 (syn. Xanthocalanus minor Giesbrecht, 1893)
  • Xanthocalanus giesbrechti Thompson I.C., 1903
  • Xanthocalanus globiceps (Farran, 1908) (syn. Amallophora globiceps (Farran, 1908)
  • Xanthocalanus gracilis Wolfenden, 1911
  • Xanthocalanus greeni Farran, 1905 (syn. Xanthocalanus greenii Farran, 1905; Talacalanus greenii (Farran, 1905)
  • Xanthocalanus groenlandicus Tupitzky, 1982 (syn.Tharybis groenlandicus (Tupitzky, 1982): Tharybis groenlandica (Tupitzky, 1982)
  • Xanthocalanus harpagatus Bradford & Wells, 1983
  • Xanthocalanus hirtipes Vanhöffen, 1897
  • Xanthocalanus hispidus Grice & Hulsemann, 1967
  • Xanthocalanus impar (Wolfenden, 1911) (syn. Scaphocalanus impar (Wolfenden, 1911)
  • Xanthocalanus incertus Sars G.O., 1920
  • Xanthocalanus irritans (Tanaka, 1960)
  • Xanthocalanus kurilensis Brodsky, 1950
  • Xanthocalanus laptevorum Markhaseva, 1998 (= Byrathis laptevorum (Markhaseva, 1998)
  • Xanthocalanus legatus Tanaka, 1960
  • Xanthocalanus longispinus Bradford-Grieve, 2004
  • Xanthocalanus macilenta (Grice & Hulsemann, 1970)
  • Xanthocalanus macrocephalon Grice & Hulsemann, 1970 (syn.Tharybis macrocephalon (Grice & Hulsemann, 1970); Byrathis macrocephalon (Grice & Hulsemann, 1970)
  • Xanthocalanus magnus Wolfenden, 1906 (= Onchocalanus magnus (Wolfenden, 1906)
  • Xanthocalanus marlyae Campaner, 1978
  • Xanthocalanus maximus Brodsky, 1950 (syn.Talacalanus maximus (Brodsky, 1950)
  • Xanthocalanus medius Tanaka, 1937
  • Xanthocalanus meteorae Markhaseva & Schnack-Schiel, 2003
  • Xanthocalanus minor Giesbrecht, 1893
  • Xanthocalanus mixtus Sars G.O., 1920
  • Xanthocalanus multispinus Chen & Zhang, 1975
  • Xanthocalanus muticus Sars G.O., 1905
  • Xanthocalanus oculatus (Tanaka, 1960)
  • Xanthocalanus obtusus Farran, 1904
  • Xanthocalanus ordinarius Grice, 1973 (= Brachycalanus ordinarius (Grice, 1973)
  • Xanthocalanus paraincertus Grice & Hulsemann, 1965 (syn. Brodskius robustipes (Grice & Hulsemann, 1965)
  • Xanthocalanus paululus Park, 1970 (= Brodskius paululus (Park, 1970)
  • Xanthocalanus pavlovskii Brodsky, 1955
  • Xanthocalanus pectinatus Tanaka, 1960
  • Xanthocalanus penicillatus Tanaka, 1960
  • Xanthocalanus pinguis Farran, 1905
  • Xanthocalanus polaris Brodsky, 1950 (=Scolecitrichopsis polaris (Brodsky, 1950)
  • Xanthocalanus polarsternae Markhaseva, 1998
  • Xanthocalanus profundus Sars G.O., 1907
  • Xanthocalanus propinquus Sars G.O., 1902
  • Xanthocalanus ptevorum Markhaseva, 1998 [Laptev Sea]
  • Xanthocalanus pulcher Esterly, 1911
  • Xanthocalanus quasiprofundus Vyshkvartzeva, 2002
  • Xanthocalanus robustus (Scott T., 1894) (syn.Amallothrix robusta (Scott T., 1894)
  • Xanthocalanus rotundus (Grice & Hulsemann, 1970)
  • Xanthocalanus serratus (Tanaka, 1960)
  • Xanthocalanus similis Esterly, 1906 (syn. Onchocalanus cristatus (Wolfenden, 1904)
  • Xanthocalanus simplex Aurivillius, 1898 (syn.Undinella simplex (Wolfenden, 1906)
  • Xanthocalanus smithae (Grice, 1962) (syn. Scopalatum smithae (Grice, 1962)
  • Xanthocalanus soaresmoreirai Björnberg T.K.S., 1975 (syn. Pseudoamallothrix soaresmoreirai (Björnberg T.K.S., 1975)
  • Xanthocalanus spinodenticulatus Markhaseva, 1998 [Laptev Sea]
  • Xanthocalanus squamatus Farran, 1936
  • Xanthocalanus stewarti Bradford-Grieve, 2004
  • Xanthocalanus subagilis Wolfenden, 1904
  • Xanthocalanus subbrevicornis (syn. Scaphocalanus subbrevicornis (Wolfenden, 1911)
  • Xanthocalanus subcristatus Wolfenden, 1906 (syn. Onchocalanus subcristatus Wolfenden, 1911
  • Xanthocalanus tectus Esterly, 1911 (syn. Cephalophanes tectus (Esterly, 1911)
  • Xanthocalanus tenuiremis Scott T., 1909
  • Xanthocalanus tenuiserratus Wolfenden, 1911
  • Xanthocalanus typicus (Scott T., 1894)
  • Xanthocalanus validus (Farran, 1908) (syn.Amallothrix valida (Farran, 1908)
  • Xanthocalanus watersae Grice, 1973 (syn. Neoscolecithrix watersae (Grice, 1973)

    Key to Megacalanidae genera




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