EPACTERISCIDAE Fosshagen, 1973
"The family Epacteriscidae was established by Fosshagen in 1973 to accommodate a new monotypic genus of
calanoid, Epacteriscus Fosshagen, 1973, taken in bottom dredges and coral rubble washings in Florida and
Colombia (Fosshagen 1973)" (Boxshall1& Jaume 2003)
"The Epacteriscidae is a relatively recently discovered
demersal family within the order Calanoida; most species
of which have been obtained from flooded coastal
caves in warm temperate to tropical shallow waters. Its
members are characterized by a bilobed rostrum bearing
filaments, up to 27 segments in the female antennules,
raptorial mouthparts and by primitive swimming
legs with 3-segmented rami" (from: Fosshagen et al., 2001) 
AZYGONECTES Fosshagen & Iliffe, 2004
Azygonectes intermedius Fosshagen & Iliffe, 2004
BALINELLA Fosshagen, Boxshall & Iliffe, 2001
Balinella ornata Fosshagen, Boxshall & Iliffe, 2001
Balinella yucatanensis Suárez-Morales, Ferrari & Iliffe, 2006BOFURIELLA Fosshagen, Boxshall & Iliffe, 2001
Bofuriella paravorata Fosshagen & Iliffe, 2007
Bofuriella spinosa Fosshagen & Iliffe, 2007
Bofuriella vorata Fosshagen, Boxshall & Iliffe, 2001
BOMBURIELLA Fosshagen, Boxshall & Iliffe, 2001
Bomburiella gigas Fosshagen, Boxshall & Iliffe, 2001
BUNDERIA Jaume & Humphreys, 2001
Bunderia misophaga Jaume & Humphreys, 2001
CAICONECTES Fosshagen & Iliffe, 2007
Caiconectes antiquus Fosshagen & Iliffe, 2007
CRYPTONECTES Fosshagen & Iliffe, 2004
Cryptonectes brachyceratus Fosshagen & Iliffe, 2004
EDAXIELLA Fosshagen, Boxshall & Iliffe, 2001
Edaxiella rubra Fosshagen, Boxshall & Iliffe, 2001
ENANTIOSIS Barr, 1984
Enantiosis belizensis Fosshagen, Boxshall & Iliffe, 2001
Enantiosis bermudensis Fosshagen, Boxshall & Iliffe, 2001
Enantiosis cavernicola Barr, 1984
Enantiosis conspinulata Fosshagen, Boxshall & Iliffe, 2001 Fosshagen, Boxshall & Iliffe, 2001
Enantiosis dicerata Fosshagen, Boxshall & Iliffe, 2001
Enantiosis dicerata Fosshagen, Boxshall & Iliffe, 2001 Fosshagen, Boxshall & Iliffe, 2001
Enantiosis galapagensis Fosshagen, Boxshall & Iliffe, 2001
Enantiosis longiprocessa Fosshagen, Boxshall & Iliffe, 2001 Fosshagen, Boxshall & Iliffe, 2001
ENANTRONIA Fosshagen, Boxshall & Iliffe, 2001
Enantronia canariensis Fosshagen, Boxshall & Iliffe, 2001
ENANTRONOIDES Fosshagen, Boxshall & Iliffe, 2001
Enantronoides bahamensis Fosshagen, Boxshall & Iliffe, 2001
EPACTERISCUS Fosshagen, 1973 Fosshagen, Boxshall & Iliffe, 2001
Epacteriscus cuspidantennula Fosshagen, Boxshall & Iliffe, 2001 Fosshagen, Boxshall & Iliffe, 2001
Epacteriscus dentipes Fosshagen, Boxshall & Iliffe, 2001 Fosshagen, Boxshall & Iliffe, 2001
Epacteriscus rapax Fosshagen, 1973 Fosshagen, Boxshall & Iliffe, 2001
EREBONECTES Fosshagen, in Fosshagen & Iliffe, 1985
Erebonectes nesioticus Fosshagen, in Fosshagen & Iliffe, 1985
Erebonectes macrochaetus Fosshagen & Iliffe, 1994 (syn. Erebonectoides macrochaetus (Fosshagen & Iliffe, 1994)
EREBONECTOIDES Fosshagen, Boxshall & Iliffe, 2001
Erebonectoides macrochaetus (Fosshagen & Iliffe, 1994)
GLOINELLA Fosshagen, Boxshall & Iliffe, 2001
Gloinella yagerae Fosshagen, Boxshall & Iliffe, 2001
IBOYELLA Boxshall & Jaume, 2003
Iboyella cubensis Boxshall & Jaume, 2003
MIHEPTNERIA
Miheptneria abyssalis Andronov, 2007
MINNONECTES Fosshagen & Iliffe, 2004
Minnonectes melodactylus Fosshagen & Iliffe, 2004
OINELLA Fosshagen, Boxshall & Iliffe, 2001
Oinella longiseta Fosshagen, Boxshall & Iliffe, 2001
KEY TO SPECIES OF EPACTERISCIDAE
EUCALANIDAE Giesbrecht, 1893
Copepods of the family Eucalanidae Giesbrecht,
1893 are an essential component of zooplankton
communities in the tropical, boreal,
and subantarctic zones of the World Ocean. Eucalanids
can achieve high abundances in oceanic
ecosystems (Arcos & Fleminger, 1986; Heinrikh,
1990; Lavaniegos & Lopez-Cortes, 1997; Suarez-
Morales, 1998), and as large-sized copepods, they
often dominate in terms of biomass (Vinogradov
& Arashkevich, 1969; Musaeva & Kolosova,
1995; Prusova & Smith, 2005) (from:Prusova & Al-Hashmi 2024.
EUCALANUS Dana, 1852-1853
Eucalanus acus Farran, 1929 syn. Eucalanus longiceps Matthews, 1925; Subeucalanus longiceps (Matthews, 1925)
Eucalanus attenuatus (Dana, 1849) syn. Pareucalanus attenuatus (Dana, 1849)
Eucalanus bungii Giesbrecht, 1893
Eucalanus californicus Johnson M.W., 1938
Eucalanus crassus Giesbrecht, 1888 syn. Subeucalanus crassus (Giesbrecht, 1888)
Eucalanus dentatus Scott A., 1909 syn. Subeucalanus dentatus (Scott A., 1909)
Eucalanus elongatus elongatus (Dana, 1848)
Eucalanus elongatus inermis Giesbrecht, 1893
Eucalanus elongatus bungii Johnson M.W., 1938 syn. Eucalanus bungii Giesbrecht, 1893
Eucalanus elongatus hyalinus Claus, 1866 syn.Eucalanus elongatus (Dana, 1848); Eucalanus elongatus elongatus (Dana, 1848)
Eucalanus hyalinus (Claus, 1866)
Eucalanus inermis Giesbrecht, 1893
Eucalanus langae Fleminger, 1973 syn. Pareucalanus langae (Fleminger, 1973)
Eucalanus longiceps Matthews, 1925 syn. Subeucalanus longiceps (Matthews, 1925)
Eucalanus monachus Giesbrecht, 1888 syn. Subeucalanus monachus (Giesbrecht, 1888)
Eucalanus mucronatus Giesbrecht, 1888 syn. Subeucalanus mucronatus (Giesbrecht, 1888)
Eucalanus oculanus Marukawa, 1921 syn. Eucalanus crassus Giesbrecht, 1888; Subeucalanus crassus (Giesbrecht, 1888)
Eucalanus muticus Wilson C.B., 1950
Eucalanus parki Fleminger, 1973 syn. Pareucalanus parki (Fleminger, 1973)
Eucalanus peruanus Volkov, 1971 syn. Pareucalanus peruanus (Volkov, 1971)
Eucalanus pileatus Giesbrecht, 1888 syn. Subeucalanus pileatus (Giesbrecht, 1888)
Eucalanus pseudattenuatus Sewell, 1947 syn. Eucalanus attenuatus (Dana, 1849); Pareucalanus attenuatus (Dana, 1849)
Eucalanus quadrisetosus Geletin, 1973 syn.Pareucalanus parki (Fleminger, 1973)
Eucalanus saravatae Oliveira, 1945 syn. Subeucalanus pileatus (Giesbrecht, 1888)
Eucalanus sewelli Fleminger, 1973 syn. Pareucalanus sewelli (Fleminger, 1973)
Eucalanus spinifer Scott T., 1894 syn. Eucalanus elongatus (Dana, 1848); Eucalanus elongatus elongatus (Dana, 1848)
Eucalanus subcrassus Giesbrecht, 1888 syn.Subeucalanus subcrassus (Giesbrecht, 1888)
Eucalanus subtenuis Giesbrecht, 1888 syn. Subeucalanus subtenuis (Giesbrecht, 1888)
Eucalanus subtenuis japonica Fukase, 1957 syn. Subeucalanus subtenuis japonicus (Fukase, 1957)
Eucalanus vadicola Dahl F., 1894 syn. Subeucalanus monachus (Giesbrecht, 1888)
Subeucalanus flemingeri Prusova et al., 2001 (?)
PAREUCALANUS Geletin, 1976
Pareucalanus attenuatus (Dana, 1849)
Pareucalanus langae (Fleminger, 1973)
Pareucalanus parki (Fleminger, 1973)
Pareucalanus peruanus (Volkov, 1971)
Pareucalanus sewelli (Fleminger, 1973)
Pareucalanus smithae Prusova, 2007
SUBEUCALANUS Geletin, 1976
Subeucalanus dextrolateralis Prusova & Al-Hashmi 2024 [Sea of Oman] Prusova & Al-Hashmi 2024
Rhincalaninae Geletin, 1976
Subeucalaninae Geletin, 1976
Calanella Claus, 1863 syn. Eucalanus Dana, 1852-1853
DISCOIDAE Gordejeva, 1975
DISCO Grice & Hulsemann, 1965
Disco atlanticus Gordeeva K.T., 1974
Disco caribbeanensis Gordeeva K.T., 1974
Disco compressus Schulz, 2003
Disco creatus Gordeeva K.T., 1975
Disco curtirostris Gordeeva K.T., 1975
Disco elephantus Gordeeva K.T., 1975
Disco erythraeus Gordeeva K.T., 1974
Disco fjordicus Fosshagen, 1967 (syn. Disco fiordicus Fosshagen, 1967; Pertsovius fjordicus (Fosshagen, 1967)
Disco fiordicus Fosshagen, 1967 (= Pertsovius fjordicus (Fosshagen, 1967)
Disco hartmanni Schulz, 1993
Disco inflatus Grice & Hulsemann, 1965
Disco intermedius Gordeeva K.T., 1976
Disco longus Grice & Hulsemann, 1965 (syn. Pertsovius longus (Grice & Hulsemann, 1965)
Disco marinus Gordeeva K.T., 1974
Disco mediterraneus Gordeeva K.T., 1974 (syn. Paradisco mediterraneus (Gordeeva K.T., 1974)
Disco minutus Grice & Hulsemann, 1965
Disco oceanicus Gordeeva K.T., 1974 (syn. Pertsovius oceanicus (Gordeeva K.T., 1974)
Disco oviformis Park, 1970 (syn. Pertsovius oviformis (Park, 1970)
Disco peltatus Gordeeva K.T., 1974
Disco populosus Gordeeva K.T., 1976
Disco robustipes Gordeeva K.T., 1974
Disco triangularis Markhaseva & Kosobokova, 1998
Disco tropicus Gordeeva K.T., 1974
Disco vulgaris Gordeeva K.T., 1974
PARADISCO Gordeeva, 1975
Paradisco gracilis Gordeeva K.T., 1975
Paradisco grandis Gordeeva K.T., 1976
Paradisco mediterraneus (Gordeeva K.T., 1974)
Paradisco nudus Schulz, 1993
PERTSOVIUS Andronov & Kosobokova, 2011
Pertsovius fjordicus (Fosshagen, 1967)
Pertsovius heterodentatus Andronov & Kosobokova, 2011
Pertsovius longus (Grice & Hulsemann, 1965)
Pertsovius oceanicus (Gordeeva K.T., 1974)
Pertsovius oviformis (Park, 1970)
Pertsovius serratus Andronov & Kosobokova, 2011
Pertsovius tridentatus Andronov & Kosobokova, 2011
PRODISCO Gordeeva, 1975
Prodisco princeps Gordeeva K.T., 1975
Prodisco secundus Gordeeva K.T., 1975
NULLOSETIGERIDAE Soh, Ohtsuka, Imabayashi & Suh, 1999
This family contains only one genus Nullosetigera Soh, Ohtsuka, Imabayashi, Suh, 1999. This genus was previously known as Phyllopus Brady, 1883 but this name was discovered to be a preoccupied (Soh et al. 1999).
NULLOSETIGERA Soh, Ohtsuka, Imabayashi & Suh, 1999 (syn. Phyllopus Brady, 1883)
Nullosetigera aequalis (Sars G.O., 1920)
Nullosetigera auctiseta Soh, Ohtsuka, Imabayashi & Suh, 1999
Nullosetigera bidentata (Brady, 1883)
Nullosetigera giesbrechti (Scott A., 1909)
Nullosetigera helgae (Farran, 1908)
Nullosetigera impar (Farran, 1908)
Nullosetigera integer (Esterly, 1911)
Nullosetigera mutata (Tanaka, 1964)
Nullosetigera mutica (Sars G.O., 1907)
Nullosetigera turqueti (Quidor, 1906)
SPINOCALANIDAE Vervoort, 1951
DAMKAERIA Fosshagen, 1983
Damkaeria bicornuta Schulz, 2006
Damkaeria falcifera Fosshagen, 1983
FOXTONIA Hulsemann & Grice, 1963 (Family Arctoconstantinidae Markhaseva & Kosobokova, 2001)
Foxtonia barbatula Hulsemann & Grice, 1963
ISAACSICALANUS Fleminger, 1983
Isaacsicalanus paucisetus Fleminger, 1983
KUNIHULSEA Schulz, 1992
Kunihulsea antarctica Schulz, 2004
Kunihulsea arabica Schulz, 1992
METHANOCALANUS Ivanenko, Defaye & Cuoc, 2007
Methanocalanus gabonicus Ivanenko, Defaye & Cuoc, 2007
MIMOCALANUS Farran, 1908
Mimocalanus brodskyi Razouls, 1974
Mimocalanus crassus Park, 1970
Mimocalanus cultrifer Farran, 1908
Mimocalanus damkaeri Brodsky, Vyshkvartseva, Kos & Markhaseva, 1983
Mimocalanus distinctocephalus Brodsky, 1950(= Mimocalanus nudus Farran, 1908)
Mimocalanus heronae Damkaer, 1975
Mimocalanus inflatus Davis, 1949
Mimocalanus major Sars G.O., 1920
Mimocalanus nudus Farran, 1908
Mimocalanus ovalis (Grice & Hulsemann, 1965)
Mimocalanus sulcifrons Wheeler, 1970 (=Mimocalanus nudus Farran, 1908)
MONACILLA Sars G.O., 1905 (syn. Hypsicalanus Wolfenden, 1911;Oxycalanus Farran, 1908)
Monacilla dubia Scott A., 1909 (syn. Monacilla typica Sars G.O., 1905)
Monacilla gracilis (Wolfenden, 1911)
Monacilla semispina (Scott A., 1909) (syn. Monacilla typica Sars G.O., 1905)
Monacilla spinifer (Farran, 1908) (syn.Monacilla typica Sars G.O., 1905)
Monacilla tenera Sars G.O., 1907
Monacilla typica Sars G.O., 1905
MOSPICALANUS Schulz, 1996
Mospicalanus schielae Schulz, 1996
RHINOMAXILLARIS Grice & Hulsemann, 1967
Rhinomaxillaris bathybia Grice & Hulsemann, 1967 SOGNOCALANUS Fosshagen, 1967 (Family Arctoconstantinidae Markhaseva & Kosobokova, 2001)
Sognocalanus confertus Fosshagen, 1967
SPINOCALANUS Giesbrecht, 1888
Spinocalanus abruptus Grice & Hulsemann, 1965
Spinocalanus abyssalis Giesbrecht, 1888
Spinocalanus angusticeps Sars G.O., 1920
Spinocalanus antarcticus Wolfenden, 1906
Spinocalanus aspinosus Park, 1970
Spinocalanus brevicaudatus Brodsky, 1950
Spinocalanus caudatus Sars G.O., 1920
Spinocalanus dispar Schulz, 1989
Spinocalanus dorsispinosus Brodsky, 1950(syn. Spinocalanus horridus Wolfenden, 1911)
Spinocalanus elongatus Brodsky, 1950
Spinocalanus giesbrechti Brady, 1918 (taxon inquirendum)
Spinocalanus heterocaudatus Rose, 1937 (syn. Spinocalanus magnus Wolfenden, 1904)
Spinocalanus hirtus Sars G.O., 1907
Spinocalanus hoplites Park, 1970
Spinocalanus horridus Wolfenden, 1911
Spinocalanus latifrons Sars G.O., 1907( syn. Spinocalanus magnus Wolfenden, 1904)
Spinocalanus longicornis Sars G.O., 1900
Spinocalanus longipes Tanaka, 1956 (syn. Spinocalanus angusticeps Sars G.O., 1920)
Spinocalanus longispinus Brodsky, 1950 (syn. Spinocalanus horridus Wolfenden, 1911)
Spinocalanus macrocephalon Brodsky, Vyshkvartseva, Kos & Markhaseva, 1983
Spinocalanus magnus Wolfenden, 1904
Spinocalanus major Esterly, 1906 (syn.Spinocalanus brevicaudatus Brodsky, 1950)
Spinocalanus neospinosus Grice, 1971 (syn. Spinocalanus oligospinosus Park, 1970)
Spinocalanus oligospinosus Park, 1970
Spinocalanus ovalis Grice & Hulsemann, 1965 (syn.Mimocalanus ovalis (Grice & Hulsemann, 1965)
Spinocalanus pacificus Mori, 1942 (syn. Spinocalanus magnus Wolfenden, 1904)
Spinocalanus parabyssalis Park, 1970 (syn. Spinocalanus longicornis Sars G.O., 1900)
Spinocalanus paraoligospinosus Hure & Scotto di Carlo, 1971 (nomen nudum)
Spinocalanus pteronus Park, 1970 (syn. Spinocalanus usitatus Park, 1970)
Spinocalanus polaris Brodsky, 1950
Spinocalanus profundalis stat.nov. (?)
Spinocalanus pseudospinipes Brodsky, 1950 (syn. Spinocalanus brevicaudatus Brodsky, 1950)
Spinocalanus pteronus Park, 1970
Spinocalanus schaudinni Mrázek, 1902 (syn. Spinocalanus longicornis Sars G.O., 1900)
Spinocalanus similis Brodsky, 1950
Spinocalanus spinipes Brodsky, 1950 (syn.Spinocalanus horridus Wolfenden, 1911)
Spinocalanus spinosus Farran, 1908
Spinocalanus stellatus Brodsky, 1950 (syn.Spinocalanus horridus Wolfenden, 1911)
Spinocalanus terranovae Damkaer, 1975
Spinocalanus usitatus Park, 1970
Spinocalanus validus Sars G.O., 1920
Spinocalanus ventriosus Grice & Hulsemann, 1967 (syn.Mimocalanus ovalis (Grice & Hulsemann, 1965)
KEY TO SPECIES OF SPINOCALANUS
TENERIFORMA Grice & Hulsemann, 1967 (syn.Hypsicalanus Wolfenden, 1911; Monacilla Sars G.O., 1905;
Oxycalanus Farran, 1908 ayn. Monacilla Sars G.O., 1905;
Tanyrhinus Farran, 1936 syn. Teneriforma Grice & Hulsemann, 1967
Teneriforma meteorae Schulz, 1989
Teneriforma naso (Farran, 1936)
Teneriforma pakae Andronov, 2013
Teneriforma pentatrichodes Schulz, 1989
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