HETERORHABDIDAE Sars G.O., 1902

This family consists of 67 species in eight genera worldwide (Walter and Boxshall, 2020)

DISSETA Giesbrecht, 1889

  • Disseta atlantica Wolfenden, 1911 (syn. Disseta palumbii Giesbrecht, 1889
  • Disseta coelebs Heptner, 1972 (syn. Disseta magna Bradford, 1971
  • Disseta grandis Esterly, 1906 (syn. Disseta palumbii Giesbrecht, 1889
  • Disseta magna Bradford, 1971
  • Disseta maxima Esterly, 1911(syn. Disseta scopularis (Brady, 1883)
  • Disseta palumbii Giesbrecht, 1889
  • Disseta minuta Grice & Hulsemann, 1965 (syn. Microdisseta minuta (Grice & Hulsemann, 1965)
  • Disseta scopularis (Brady, 1883)

    HEMIRHABDUS Wolfenden, 1911 (syn. Hemirhabus Wolfenden, 1911; Macrorhabdus Sars G.O., 1920)

  • Hemirhabdus amplus Park, 2000
  • Hemirhabdus falciformis Wolfenden, 1911 (syn. Neorhabdus falciformis (Wolfenden, 1911)
  • Hemirhabdus grimaldii (Richard, 1893)
  • Hemirhabdus latus (Sars G.O., 1905) (syn. Neorhabdus latus (Sars G.O., 1905)
  • Hemirhabdus truncatus (Scott A., 1909) (syn. Neorhabdus truncatus (Scott A., 1909)

    HETERORHABDUS Giesbrecht, 1898 (syn. Alloiorhabdus Wolfenden, 1911; Heterochaeta Claus, 1863)

  • Heterorhabdus abyssalis (Giesbrecht, 1889)
  • Heterorhabdus americanus Park, 2000
  • Heterorhabdus ankylocolus Park, 2000
  • Heterorhabdus atlanticus Wolfenden, 1905
  • Heterorhabdus austrinus Giesbrecht, 1902
  • Heterorhabdus brevicornis (Dahl F., 1894)
  • Heterorhabdus caribbeanensis Park, 1970
  • Heterorhabdus clausi (Giesbrecht, 1889) (syn. Heterorhabdus clausi (Giesbrecht, 1889)
  • Heterorhabdus compactus (Sars G.O., 1900) (syn. Paraheterorhabdus compactus (Sars G.O., 1900)
  • Heterorhabdus confusibilis Park, 2000
  • Heterorhabdus compactoides Heptner, 1971 (syn. Paraheterorhabdus compactoides (Heptner, 1971)
  • Heterorhabdus compactus (Sars G.O., 1900) (syn.Paraheterorhabdus compactus (Sars G.O., 1900)
  • Heterorhabdus dubius Tanaka, 1953 (syn. Heterorhabdus abyssalis (Giesbrecht, 1889)
  • Heterorhabdus egregius Heptner, 1972
  • Heterorhabdus farrani Brady, 1918 (syn. Paraheterorhabdus farrani (Brady, 1918)
  • Heterorhabdus fistulosus Tanaka, 1964
  • Heterorhabdus grandis Wolfenden, 1904 (syn.Disseta palumbii Giesbrecht, 1889
  • Heterorhabdus grimaldii (Richard, 1893) (syn. Hemirhabdus grimaldii (Richard, 1893)
  • Heterorhabdus guineanensis Park, 2000
  • Heterorhabdus habrosomus Park, 2000
  • Heterorhabdus heterolobus Park, 2000
  • Heterorhabdus insukae Park, 2000
  • Heterorhabdus latus Sars G.O., 1905 (syn. Neorhabdus latus (Sars G.O., 1905)
  • Heterorhabdus lobatus Bradford, 1971
  • Heterorhabdus longicornis (Giesbrecht, 1889) (syn. Heterostylites longicornis (Giesbrecht, 1889)
  • Heterorhabdus longisegmentus Park, 2000
  • Heterorhabdus longispinus Davis, 1949 (syn. Paraheterorhabdus longispinus (Davis, 1949)
  • Heterorhabdus major (Dahl F., 1894) (syn. Heterostylites major (Dahl F., 1894)
  • Heterorhabdus medianus Park, 1970 (syn. Paraheterorhabdus medianus (Park, 1970)
  • Heterorhabdus medius (Wolfenden, 1911)
  • Heterorhabdus nigrotinctus Brady, 1918 (syn. Heterostylites nigrotinctus (Brady, 1918)
  • Heterorhabdus norvegicus (Boeck, 1872)
  • Heterorhabdus oikoumenikis Park, 2000
  • Heterorhabdus pacificus Brodsky, 1950
  • Heterorhabdus papilliger (Claus, 1863)
  • Heterorhabdus paraspinosus Park, 2000
  • Heterorhabdus pilosus Tanaka, 1953 (syn. Heterorhabdus tanneri (Giesbrecht, 1895)
  • Heterorhabdus profundus (Dahl F., 1894)
  • Heterorhabdus prolatus Park, 2000
  • Heterorhabdus prolixus Park, 2000
  • Heterorhabdus proximus Davis, 1949 (syn. Heterorhabdus tanneri (Giesbrecht, 1895)
  • Heterorhabdus pustulifer Farran, 1929
  • Heterorhabdus pustulifer Farran, 1929
  • Heterorhabdus quadrilobus Park, 2000
  • Heterorhabdus robustoides Brodsky, 1950 (syn. Paraheterorhabdus robustus (Farran, 1908)
  • Heterorhabdus robustus Farran, 1908 (syn. Paraheterorhabdus robustus (Farran, 1908)
  • Heterorhabdus spinifer Park, 1970
  • Heterorhabdus spinifrons (Claus, 1863)
  • Heterorhabdus spinosus Bradford, 1971
  • Heterorhabdus subspinifrons Tanaka, 1964
  • Heterorhabdus tanneri (Giesbrecht, 1895)
  • Heterorhabdus tenuis Tanaka, 1964 (syn. Paraheterorhabdus vipera (Giesbrecht, 1889)
  • Heterorhabdus tropicus (Dahl F., 1894)
  • Heterorhabdus tuberculus Park, 2000
  • Heterorhabdus vicinus Tanaka, 1964
  • Heterorhabdus viperus (Giesbrecht, 1889) (syn. Paraheterorhabdus vipera (Giesbrecht, 1889)
  • Heterorhabdus zetesios (Wolfenden, 1902) (syn. Heterostylites major (Dahl F., 1894)

    HETEROSTYLITES Sars G.O., 1920

  • Heterostylites echinatus Park, 2000
  • Heterostylites longicornis (Giesbrecht, 1889)
  • Heterostylites longioperculis Park, 2000
  • Heterostylites major (Dahl F., 1894)
  • Heterostylites nigrotinctus (Brady, 1918)
  • Heterostylites submajor Park, 2000

    MESORHABDUS Sars G.O., 1905

  • Mesorhabdus annectens Sars G.O., 1905 (syn. Mesorhabdus brevicaudatus (Wolfenden, 1905)
  • Mesorhabdus angustus Sars G.O., 1907
  • Mesorhabdus brevicaudatus (Wolfenden, 1905)
  • Mesorhabdus gracilis Sars G.O., 1907
  • Mesorhabdus paragracilis Park, 2000
  • Mesorhabdus poriphorus Park, 2000
  • Mesorhabdus truncatus Scott A., 1909 (Mesorhabdus trubeatus in the original description) (syn. Neorhabdus truncatus (Scott A., 1909)

    MICRODISSETA Heptner, 1972

  • Microdisseta minuta (Grice & Hulsemann, 1965)

    NEORHABDUS Heptner, 1972

  • Neorhabdus brevicornis Park, 2000
  • Neorhabdus capitaneus Park, 2000
  • Neorhabdus falciformis (Wolfenden, 1911)
  • Neorhabdus latus (Sars G.O., 1905)
  • Neorhabdus subcapitaneus Park, 2000
  • Neorhabdus truncatus (Scott A., 1909) (syn. Neorhabdus latus (Sars G.O., 1905)

    PARAHETERORHABDUS Brodsky, 1950

  • Paraheterorhabdus compactoides (Heptner, 1971)
  • Paraheterorhabdus farrani (Brady, 1918)
  • Paraheterorhabdus compactus (Sars G.O., 1900)
  • Paraheterorhabdus farrani (Brady, 1918)
  • Paraheterorhabdus illgi Park, 2000
  • Paraheterorhabdus longispinus (Davis, 1949)
  • Paraheterorhabdus medianus (Park, 1970)
  • Paraheterorhabdus norvegicus (Boeck, 1872) (syn. Heterorhabdus norvegicus (Boeck, 1872)
  • Paraheterorhabdus robustus (Farran, 1908)
  • Paraheterorhabdus vipera (Giesbrecht, 1889)

    Alloiorhabdus Wolfenden, 1911 accepted as Heterorhabdus Giesbrecht, 1898 (synonym); Hemirhabus Wolfenden, 1911 accepted as Hemirhabdus Wolfenden, 1911 (Lapsus calami): Heterochaeta Claus, 1863 accepted as Heterorhabdus Giesbrecht, 1898 (synonym, preoccupied name of a bird species); Macrorhabdus Sars G.O., 1920 accepted as Hemirhabdus Wolfenden, 1911.


    METRIDINIDAE Sars G.O., 1902 (syn. Metridiidae Sars G.O., 1902)

    GAUSSIA Wolfenden, 1905

  • Gaussia asymmetrica Björnberg T.K.S. & Campaner, 1988
  • Gaussia intermedia Defaye, 1998
  • Gaussia melanotica Wolfenden, 1905
  • Gaussia princeps (Scott T., 1894)
  • Gaussia scotti (Giesbrecht, 1897) (syn. Gaussia princeps (Scott T., 1894)
  • Gaussia sewelli Saraswathy, 1973

    METRIDIA Boeck, 1865 syn. Metridina Norman, 1878)

  • Metridia alata Roe, 1975
  • Metridia andraeana Brady, 1918
  • Metridia armata Boeck, 1865 (syn. Metridia longa (Lubbock, 1854)
  • Metridia asymmetrica Brodsky, 1950
  • Metridia atra Esterly, 1906 (syn. Metridia princeps Giesbrecht, 1889
  • Metridia bicornuta Davis, 1949
  • Metridia boeckii Giesbrecht, 1889
  • Metridia brevicauda Giesbrecht, 1889
  • Metridia calypsoi Gaudy, 1963
  • Metridia curticauda Giesbrecht, 1889
  • Metridia discreta Farran, 1947
  • Metridia effusa Grice & Hulsemann, 1967
  • Metridia ferrarii Markhaseva, 2001
  • Metridia gerlachei Giesbrecht, 1902
  • Metridia gurjanovae Epstein, 1949
  • Metridia hibernica (Brady & Robertson, 1873) (syn. Metridia lucens Boeck, 1865)
  • Metridia ignota Esterly, 1906
  • Metridia longa (Lubbock, 1854)
  • Metridia lucens lucens Boeck, 1865
  • Metridia macrura Sars G.O., 1905
  • Metridia normani Giesbrecht, 1893
  • Metridia okhotensis Brodsky, 1950
  • Metridia ornata Brodsky, 1950
  • Metridia pacifica Brodsky, 1950
  • Metridia princeps Giesbrecht, 1889
  • Metridia pseudoasymmetrica Markhaseva, 2001
  • Metridia scotti Giesbrecht, 1897 (syn. Gaussia scotti (Giesbrecht, 1897)
  • Metridia similis Brodsky, 1950
  • Metridia trispinosa Brady, 1918
  • Metridia venusta Giesbrecht, 1889

    PLEUROMAMMA Giesbrecht in Giesbrecht & Schmeil, 1898 (syn. Pleuromma Claus, 1863)

  • Pleuromamma abdominalis abdominalis (Lubbock, 1856)
  • Pleuromamma antarctica Steuer, 1931
  • Pleuromamma armata Pouchet & Guerne, 1887
  • Pleuromamma borealis Dahl F., 1893
  • Pleuromamma gracilis gracilis Claus, 1863
  • Pleuromamma indica Wolfenden, 1905
  • Pleuromamma johnsoni Ferrari F.D. & Saltzman, 1998
  • Pleuromamma piseki Farran, 1929
  • Pleuromamma princeps (Scott T., 1894) (syn. Gaussia princeps (Scott T., 1894)
  • Pleuromamma quadrungulata quadrungulata (Dahl F., 1893)
  • Pleuromamma robusta robusta (Dahl F., 1893)
  • Pleuromamma scutullata Brodsky, 1950
  • Pleuromamma wolfendeni Brady, 1918
  • Pleuromamma xiphias (Giesbrecht, 1889)


    Metridina Norman, 1878 syn, Metridia Boeck, 1865; Pleuromma Claus, 1863 syn.Pleuromamma Giesbrecht in Giesbrecht & Schmeil, 1898



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