AUGAPTILIDAE Sars G.O., 1905


HALOPTILUS Giesbrecht, 1898 (syn. Hemicalanus Dana, 1852)

  • Haloptilus aculeatus (Brady, 1883)
  • Haloptilus acutifrons (Giesbrecht, 1893)
  • Haloptilus angusticeps Sars G.O., 1907
  • Haloptilus austini Grice, 1959
  • Haloptilus bulliceps Farran, 1926
  • Haloptilus caribbeanensis Park, 1970
  • Haloptilus chierchiae (Giesbrecht, 1889)
  • Haloptilus fertilis (Giesbrecht, 1893)
  • Haloptilus fons Farran, 1908
  • Haloptilus furcatus Sars G.O., 1920
  • Haloptilus longiceps Tanaka, 1964(= Haloptilus austini Grice, 1959
  • Haloptilus longicirrus Brodsky, 1950
  • Haloptilus longicornis (Claus, 1863)
  • Haloptilus longimanus Wolfenden, 1906 (= Pseudhaloptilus longimanus (Wolfenden, 1906)
  • Haloptilus major Wolfenden, 1911
  • Haloptilus mucronatus (Claus, 1863)
  • Haloptilus ocellatus Wolfenden, 1905
  • Haloptilus orientalis (Brady, 1883)
  • Haloptilus ornatus (Giesbrecht, 1893)
  • Haloptilus oxycephalus (Giesbrecht, 1889)
  • Haloptilus pacificus Chiba, 1956
  • Haloptilus paralongicirrus Park, 1970
  • Haloptilus plumosus (Claus, 1863)
  • Haloptilus pseudooxycephalus Brodsky, 1950
  • Haloptilus setuliger Tanaka, 1964 (= Haloptilus longicirrus Brodsky, 1950
  • Haloptilus spiniceps (Giesbrecht, 1893)
  • Haloptilus spinifrons (Sars G.O., 1900) (= Haloptilus acutifrons (Giesbrecht, 1893)
  • Haloptilus tenuis Farran, 1908
  • Haloptilus validus Sars G.O., 1920

    HETEROPTILUS Sars G.O., 1920

  • Heteroptilus acutilobus (Sars G.O., 1905)
  • Heteroptilus attenuatus (Sars G.O., 1905)

    PACHYPTILUS Sars G.O., 1920

  • Pachyptilus abbreviatus (Sars G.O., 1905) (syn. Pseudhaloptilus abbreviatus (Sars G.O., 1905)
  • Pachyptilus eurygnathus Sars G.O., 1920 (syn.Pseudhaloptilus eurygnathus (Sars G.O., 1920)
  • Pachyptilus lobatus Sars G.O., 1920
  • Pachyptilus pacificus Johnson M.W., 1936 (syn. Pseudhaloptilus pacificus (Johnson, M.W., 1936)

    PONTOPTILUS Sars G.O., 1905

  • Pontoptilus abbreviatus Sars G.O., 1905 (syn. Pachyptilus abbreviatus (Sars G.O., 1905)
  • Pontoptilus acutilobus Sars G.O., 1905 (syn. Heteroptilus acutilobus (Sars G.O., 1905)
  • Pontoptilus attenuatus Sars G.O., 1905 (syn. Heteroptilus attenuatus (Sars G.O., 1905)
  • Pontoptilus lacertosus Grice & Hulsemann, 1967
  • Pontoptilus mucronatus Sars G.O., 1905
  • Pontoptilus muticus Sars G.O., 1905
  • Pontoptilus ovalis Sars G.O., 1905
  • Pontoptilus pertenuis Sars G.O., 1905
  • Pontoptilus robustus Sars G.O., 1905

    PSEUDAUGAPTILUS Sars G.O., 1907

  • Pseudaugaptilus longiremis Sars G.O., 1907
  • Pseudaugaptilus orientalis Tanaka, 1964
  • Pseudaugaptilus polaris Brodsky, 1950

    PSEUDHALOPTILUS Wolfenden, 1911 (syn. Pseudohaloptilus Wolfenden, 1911)

  • Pseudhaloptilus abbreviatus (Sars G.O., 1905)
  • Pseudhaloptilus eurygnathus (Sars G.O., 1920)
  • Pseudhaloptilus longimanus (Wolfenden, 1906)
  • Pseudhaloptilus pacificus (Johnson, M.W., 1936)


    Hemicalanus Dana, 1852 syn.Haloptilus Giesbrecht, 1898; Neoaugaptilus Brodsky, 1950 syn.Euaugaptilus Sars G.O., 1920; Pseudohaloptilus Wolfenden, 1911 syn. Pseudhaloptilus Wolfenden, 1911.




    PARACALANIDAE Giesbrecht, 1893 (syn. Calocalanidae Bernard, 1958; Mecynoceridae Andronov, 1973)

    Dolichocerea Bernard, 1963 syn.Calocalanus Giesbrecht, 1888; Andronovius Özdikmen, 2008 syn. Delibus Andronov in Vives & Shmeleva, 2007; Bestiola Andronov, 1972 syn. Bestiolina Andronov, 1991; Delius Andronov, 1972 syn. Delibus Andronov in Vives & Shmeleva, 2007; Dolichocera Bernard, 1958 syn. Calocalanus Giesbrecht, 1888; Ischnocalanus Bernard, 1963 syn.Calocalanus Giesbrecht, 1888; Leptocalanus Bernard, 1958 syn. Calocalanus Giesbrecht, 1888; Piezocalanus Grandori, 1912 syn. Paracalanus Boeck, 1865.

    ACROCALANUS Giesbrecht, 1888

  • Acrocalanus andersoni Bowman, 1958
  • Acrocalanus gibber Giesbrecht, 1888
  • Acrocalanus gracilis Giesbrecht, 1888
  • Acrocalanus indicus Tanaka, 1960 (= Delibus nudus (Sewell, 1929)
  • Acrocalanus inermis Sewell, 1912 (= Bestiolina inermis (Sewell, 1912)
  • Acrocalanus longicornis Giesbrecht, 1888
  • Acrocalanus monachus Giesbrecht, 1888
  • Acrocalanus pediger Cleve, 1901 (=Paracalanus clevei Carl, 1907)
  • Acrocalanus similis Sewell, 1914 (=Bestiolina similis (Sewell, 1914)
  • Acrocalanus sinicus Shen & Lee, 1966 (= Bestiolina sinica (Shen & Lee, 1966)

    BESTIOLINA Andronov, 1991(syn. Bestiola Andronov, 1972)

  • Bestiolina amoyensis (Li & Huang, 1984)
  • Bestiolina arabica Ali, Al-Yamani & Prusova, 2007
  • Bestiolina brasiliensis Vieira-Menezes. Dias, Cornils, Silva & Bonecker, 2021
  • Bestiolina coreana Moon, Lee & Soh, 2010
  • Bestiolina inermis (Sewell, 1912)
  • Bestiolina mexicana Suárez-Morales & Almeyda-Artigas, 2016
  • Bestiolina okinawae Vedwe & Sakagushi, 2021 [Japan]
  • Bestiolina sarae Dorado-Roncancio & Gaviria, 2019 in: Dorado-Roncancio, Gaviria, Bernal-De La Torre & Ahrens, 2019
  • Bestiolina similis (Sewell, 1914)
  • Bestiolina sinica (Shen & Lee, 1966)
  • Bestiolina zeylonica (Andronov, 1972)

    DELIBUS Andronov in Vives & Shmeleva, 2007 (syn. Andronovius Özdikmen, 2008; Delius Andronov, 1972

  • Delibus nudus (Sewell, 1929)
  • Delibus sewelli (Björnberg T.K.S., 1982)

    MECYNOCERA Thompson I.C., 1888

  • Mecynocera clausi Thompson I.C., 1888
  • Mecynocera filicornis (Giesbrecht, 1888) (syn.Mecynocera clausi Thompson I.C., 1888)
  • Mecynocera gracilis (Tanaka, 1956) (?)
  • Mecynocera plumulosa (Claus, 1863)(syn. Calocalanus plumulosus (Claus, 1863)
  • Mecynocera tenuis (Farran, 1926)(syn. Calocalanus tenuis Farran, 1926

    PARACALANUS Boeck, 1865 (syn. Piezocalanus Grandori, 1912)

  • Paracalanus aculeatus Giesbrecht, 1888 (syn.Paracalanus aculeatus aculeatus Giesbrecht, 1888)
  • Paracalanus arabiensis Kesarkar & Anil, 2010 (syn. Parvocalanus arabiensis (Kesarkar & Anil, 2010)
  • Paracalanus brasiliensis Vieira-Menezes. Dias, Cornils, Silva & Bonecker, 2021
  • Paracalanus campaneri Björnberg T.K.S., 1982
  • Paracalanus clevei Carl, 1907 (syn. Paracalanus aculeatus Giesbrecht, 1888)
  • Paracalanus crassipes Apstein, 1912 (syn. Paracalanus crassirostris Dahl F., 1894)
  • Paracalanus crassirostris Dahl F., 1894 (sy. Parvocalanus crassirostris (Dahl F., 1894)
  • Paracalanus crassirostris Dahl F., 1894 var nudus Davis 1944
  • Paracalanus denudatus Sewell, 1929
  • Paracalanus dubia Sewell, 1912 (syn. Parvocalanus dubia (Sewell, 1912)
  • Paracalanus elegans (Andronov, 1972)(syn. Parvocalanus elegans Andronov, 1972)
  • Paracalanus elongatus (Dana, 1848) (syn. Eucalanus elongatus (Dana, 1848) r
  • Paracalanus gracilis Chen & Zhang, 1965
  • Paracalanus hibernicus Brady & Robertson, 1873 (syn. Metridia hibernica (Brady & Robertson, 1873)
  • Paracalanus indicus Wolfenden, 1905
  • Paracalanus intermedius Shen & Bai, 1956
  • Paracalanus lagunaris (Grandori, 1912) (syn. Paracalanus parvus (Claus, 1863)
  • Paracalanus latus (Andronov, 1972) (syn. Parvocalanus latus Andronov, 1972)
  • Paracalanus nanus Sars G.O., 1925
  • Paracalanus nudus Sewell, 1929 (syn.Delibus nudus (Sewell, 1929) (synonym)
  • Paracalanus orientalis Ueda et al., 2021
  • Paracalanus parvus parvus (Claus, 1863)
  • Paracalanus quasimodo Bowman, 1971
  • Paracalanus scotti Fruchtl, 1923 (syn. Parvocalanus scotti (Früchtl, 1923)
  • Paracalanus serrulus Shen & Lee, 1963
  • Paracalanus serratipes Sewell, 1912 (syn. Parvocalanus serratipes (Sewell, 1912)
  • Paracalanus tropicus Andronov, 1977

    PARVOCALANUS Andronov, 1970 (syn. Pseudoparacalanus Robinson, 1948)

  • Parvocalanus arabiensis (Kesarkar & Anil, 2010)
  • Parvocalanus brevispinatus (Shen & Lee, 1966) (syn.Parvocalanus crassirostris (Dahl F., 1894)
  • Parvocalanus crassirostris (Dahl F., 1894)
  • Parvocalanus dubia (Sewell, 1912)
  • Parvocalanus elegans Andronov, 1972
  • Parvocalanus latus Andronov, 1972
  • Parvocalanus leei Moon, Youn & Soh, 2014
  • Parvocalanus scotti (Früchtl, 1923)
  • Parvocalanus serratipes (Sewell, 1912)

    CALOCALANUS Giesbrecht, 1888 (syn. Dolichocerea Bernard, 1963; Ischnocalanus Bernard, 1963; Leptocalanus Bernard, 1958)

  • Calocalanus aculeatus Shmeleva, 1987
  • Calocalanus adriaticus Shmeleva, 1973
  • Calocalanus africanus Shmeleva, 1979
  • Calocalanus alboranus Shmeleva, 1979
  • Calocalanus antarcticus Shmeleva, 1978
  • Calocalanus atlanticus Shmeleva, 1975
  • Calocalanus beklemishevi Shmeleva, 1987
  • Calocalanus contractus Farran, 1926
  • Calocalanus curtus Andronov, 1973 syn. Calocalanus kristalli Shmeleva, 1968 (synonym according to Andronov, 1991)
  • Calocalanus dellacrocei Shmeleva, 1987
  • Calocalanus elegans Shmeleva, 1965
  • Calocalanus elongatus Shmeleva, 1968
  • Calocalanus equalicauda (Bernard, 1958)
  • Calocalanus fiolenti Shmeleva, 1978
  • Calocalanus fusiformis Shmeleva, 1978
  • Calocalanus gracilis Tanaka, 1956 syn. Mecynocera gracilis (Tanaka, 1956)
  • Calocalanus gresei Shmeleva, 1973
  • Calocalanus indicus Shmeleva, 1974
  • Calocalanus kristalli Shmeleva, 1968
  • Calocalanus latus Shmeleva, 1968
  • Calocalanus lomonosovi Shmeleva, 1975
  • Calocalanus longifurca Shmeleva, 1975
  • Calocalanus longisetosus Shmeleva, 1965
  • Calocalanus longipinus Shmeleva, 1978
  • Calocalanus minor Shmeleva, 1980
  • Calocalanus minutus Andronov, 1973
  • Calocalanus monospinus Chen & Shen, 1974
  • Calocalanus nambiensis Andronov, 1973 = Calocalanus namibiensis Andronov, 1973
  • Calocalanus namibiensis
  • Calocalanus nanus Shmeleva, 1987
  • Calocalanus neptunus Shmeleva, 1975
  • Calocalanus omaniensis Shmeleva, 1975
  • Calocalanus ovalis Shmeleva, 1965
  • Calocalanus paracontractus Shmeleva, 1974
  • Calocalanus parelongatus Shmeleva, 1979
  • Calocalanus pavo (Dana, 1848)
  • Calocalanus pavoninus Farran, 1936
  • Calocalanus plumatus Shmeleva, 1965
  • Calocalanus plumulosus (Claus, 1873)
  • Calocalanus pseudocontractus Bernard, 1958
  • Calocalanus pubes Andronov, 1973
  • Calocalanus pyriformis Shmeleva, 1975
  • Calocalanus regini Shmeleva, 1987
  • Calocalanus sayademalja Shmeleva, 1987
  • Calocalanus spinosus Shmeleva, 1987
  • Calocalanus styliremis Giesbrecht, 1888
  • Calocalanus tenuiculus Andronov, 1973
  • Calocalanus tenuis Farran, 1926
  • Calocalanus vinogradovi Shmeleva, 1987
  • Calocalanus vitjazi Shmeleva, 1974
  • Calocalanus vivesei Shmeleva, 1979
  • Calocalanus vivesi Shmeleva, 1979 (= Calocalanus vivesei Shmeleva, 1979)


    Key to the genera of family Paracalanidae




    RHINCALANIDAE Geletin, 1976

    RHINCALANUS Dana, 1852 (syn. Rhinocalanus)

  • Rhincalanus aculeatus Scott T., 1894 (syn. Arietellus aculeatus (Scott T., 1894)
  • Rhincalanus cornutus (Dana, 1849) (syn.Rhincalanus cornutus cornutus (Dana, 1849)
  • Rhincalanus gigas Brady, 1883
  • Rhincalanus grandis Giesbrecht, 1902 (syn. Rhincalanus gigas Brady, 1883
  • Rhincalanus nassatus Giesbrecht, 1888 (syn. Rhincalanus nasutus Giesbrecht, 1888
  • Rhincalanus nasutus Giesbrecht, 1888
  • Rhincalanus rostrifrons (Dana, 1849)




    TORTANIDAE Sars G.O., 1902

    TORTANUS Giesbrecht in Giesbrecht & Schmeil, 1898 (syn. Corynura Brady, 1883; Tortanus Giesbrecht in Giesbrecht & Schmeil, 1898)

  • Tortanus (Acutanus) Ohtsuka, 1992
  • Tortanus (Acutanus) angularis Ohtsuka, 1992
  • Tortanus (Acutanus) compernis Gonzalez & Bowman, 1965
  • Tortanus (Acutanus) ecornatus Ohtsuka & Reid, 1998
  • Tortanus (Acutanus) setacaudatus Williams, 1906
  • Tortanus (Atortus) ampliramus Ohtsuka, El-Sherbiny & Ueda, 2000
  • Tortanus (Atortus) andamanensis Nishida, Anandavelu & Padmavati, 2015
  • Tortanus (Atortus) bilobus Mulyadi, Nishida & Ohtsuka, 2017
  • Tortanus (Atortus) bonjoi Othman, 1987
  • Tortanus (Atortus) bowmani Othman, 1987
  • Tortanus (Atortus) brevipes Scott A., 1909
  • Tortanus (Atortus) capensis Grindley, 1978
  • Tortanus (Atortus) dhritiae Francis et al., 2022[India]
  • Tortanus (Atortus) digitalis Ohtsuka & Kimoto, 1989
  • Tortanus (Atortus) erabuensis Ohtsuka, Fukuura & Go, 1987
  • Tortanus (Atortus) giesbrechti Jones E.C. & Park, 1968
  • Tortanus (Atortus) indonesiensis Mulyadi, Nishida & Ohtsuka, 2017
  • Tortanus (Atortus) insularis Ohtsuka & Conway, 2003
  • Tortanus (Atortus) longipes Brodsky, 1948
  • Tortanus (Atortus) lophus Bowman, 1971
  • Tortanus (Atortus) lukmani Mulyadi, Nishida & Ohtsuka, 2017
  • Tortanus (Atortus) magnonyx Ohtsuka & Conway, 2005
  • Tortanus (Atortus) manadoensis Mulyadi, Nishida & Ohtsuka, 2017
  • Tortanus (Atortus) minicoyensis Francis & Bijoy Nandan, 2019
  • Tortanus (Atortus) murrayi Scott A., 1909
  • Tortanus (Atortus) nishidai Ohtsuka, El-Sherbiny & Ueda, 2000
  • Tortanus (Atortus) omorii Mulyadi, Nishida & Ohtsuka, 2017
  • Tortanus (Atortus) processus Mulyadi, Nishida & Ohtsuka, 2017
  • Tortanus (Atortus) recticaudus (Giesbrecht, 1889)
  • Tortanus (Atortus) rubidus Tanaka, 1965
  • Tortanus (Atortus) ryukyuensis Ohtsuka & Kimoto, 1989
  • Tortanus (Atortus) scaphus Bowman, 1971
  • Tortanus (Atortus) sinicus Chen, 1983
  • Tortanus (Atortus) sulawesiensis Mulyadi, Nishida & Ohtsuka, 2017
  • Tortanus (Atortus) taiwanicus Chen Q.C. & Hwang J.S., 1999
  • Tortanus (Atortus) tropicus Sewell, 1932
  • Tortanus (Atortus) tumidus Chen, Hwang & Yin, 2004
  • Tortanus (Atortus) vietnamicus Nishida & Cho, 2005
  • Tortanus (Boreotortanus) Ohtsuka & Reid, 1998
  • Tortanus (Boreotortanus) discaudatus (Thompson I.C. & Scott A.) In Herdman, Thompson & Scott, 1897)
  • Tortanus (Eutortanus) Smirnov, 1935
  • Tortanus (Eutortanus) derjugini Smirnov, 1935
  • Tortanus (Eutortanus) dextrilobatus Chen & Zhang, 1965
  • Tortanus (Eutortanus) komachi Itoh, Ohtsuka & Sato, 2001
  • Tortanus (Eutortanus) sheni Hulsemann, 1988
  • Tortanus (Eutortanus) spinicaudatus Shen & Bai, 1956
  • Tortanus (Eutortanus) terminalis Ohtsuka & Reid, 1998
  • Tortanus (Eutortanus) vermiculus Shen, 1955
  • Tortanus (Tortanus) Giesbrecht in Giesbrecht & Schmeil, 1898
  • Tortanus (Tortanus) barbatus (Brady, 1883)
  • Tortanus (Tortanus) forcipatus (Giesbrecht, 1889)
  • Tortanus (Tortanus) gracilis (Brady, 1883)
  • Tortanus (Atortus) nishidai Ohtsuka, El-Sherbiny & Ueda, 2000

    Key to the species of genus Tortanus

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