LAMIPPIDAE Joliet, 1882


ENALCYONIUM Olsson, 1869 (syn. Alcyonicola Scott T. & Scott A., 1895 )

  • Enalcyonium affinis (Zulueta, 1908)
  • Enalcyonium albidum (Zulueta, 1908)
  • Enalcyonium alcyonii (Joliet, 1882)
  • Enalcyonium auriculatum Kim I.H., 2004
  • Enalcyonium bullatum Kim I.H., 2004
  • Enalcyonium caledonensis Kim I.H., 2004
  • Enalcyonium capillatum Kim I.H., 2004
  • Enalcyonium carrikeri Dudley, 1973
  • Enalcyonium ceramensis Kim I.H., 2007
  • Enalcyonium chattoni (Zulueta, 1908) (syn. Lamippula chattoni (Zulueta, 1908)
  • Enalcyonium ciliatum Stock, 1972
  • Enalcyonium circulatum Kim I.H., 2007
  • Enalcyonium concinnum (Humes, 1957)
  • Enalcyonium confusum Stock, 1988
  • Enalcyonium digitigerum Ho, 1984
  • Enalcyonium duthiersi (Joliet, 1882) (syn. Lamippula duthiersi (Joliet, 1882)
  • Enalcyonium euniceae Stock, 1973
  • Enalcyonium forbesi (Scott T., 1901)
  • Enalcyonium grandisetigerum Kim I.H., 2009
  • Enalcyonium heegaardi Bouligand, 1960
  • Enalcyonium kohsiangi Uyeno, 2015
  • Enalcyonium humesi Kim I.H., 2004
  • Enalcyonium lobophyti Kim I.H., 2004
  • Enalcyonium nudum Stock, 1973
  • Enalcyonium olssoni (Zulueta, 1908)
  • Enalcyonium pallidum (Zulueta, 1908) (syn. Lamippula pallida (Zulueta, 1908)
  • Enalcyonium parvum (Zulueta, 1908) (syn. Lamippula parva (Zulueta, 1908)
  • Enalcyonium pusillum (Zulueta, 1908)
  • Enalcyonium ramosum Stock, 1973
  • Enalcyonium robustum Kim I.H., 2009
  • Enalcyonium rubicundum Olsson, 1869
  • Enalcyonium scorpio Stock, 1973
  • Enalcyonium setigerum (Zulueta, 1908)
  • Enalcyonium sympodii (Zulueta, 1910)
  • Enalcyonium variicauda Stock, 1973

    GORGONOPHILUS Buhl-Mortensen & Mortensen, 2004

  • Gorgonophilus canadensis Buhl-Mortensen & Mortensen, 2004

    ISIDICOLA Gravier, 1914

  • Isidicola antarctica Gravier, 1914

    LAMIPPE Bruzelius, 1858

  • Lamippe aciculifera Zulueta, 1908 (syn. Lamippina aciculifera (Zulueta, 1908)
  • Lamippe affinis Zulueta, 1908 (syn. Enalcyonium affinis (Zulueta, 1908)
  • Lamippe albida Zulueta, 1908 (syn. Enalcyonium albidum (Zulueta, 1908)
  • Lamippe alcyonii Joliet, 1882 (syn. Enalcyonium alcyonii (Joliet, 1882)
  • Lamippe anthoptili Jungersen, 1904
  • Lamippe bouligandi Laubier, 1972
  • Lamippe brementi Zulueta, 1910 (syn.Lamippina aciculifera (Zulueta, 1908)
  • Lamippe chattoni Zulueta, 1908 (syn. Lamippula chattoni (Zulueta, 1908)
  • Lamippe concinna Humes, 1957 (syn. Enalcyonium concinnum (Humes, 1957)
  • Lamippe duthiersi Joliet, 1882 (syn. Lamippula duthiersi (Joliet, 1882)
  • Lamippe faurei (Bouligand & Delamare Deboutteville, 1959) (syn. Lamippella faurei Bouligand & Delamare Deboutteville, 1959
  • Lamippe forbesi Scott T., 1901 (syn. Enalcyonium forbesi (Scott T., 1901)
  • Lamippe fusiformis (Scott T. & Scott A., 1895) (syn. Enalcyonium rubicundum Olsson, 1869
  • Lamippe olssoni Zulueta, 1908 (syn. Enalcyonium olssoni (Zulueta, 1908)
  • Lamippe pallida Zulueta, 1908 (syn. Lamippula pallida (Zulueta, 1908)
  • Lamippe papillifera Zulueta, 1910 (syn. Lamippina aciculifera (Zulueta, 1908)
  • Lamippe parva Zulueta, 1908 accepted as Lamippula parva (Zulueta, 1908)
  • Lamippe proteus Claparède, 1867
  • Lamippe pteroidis Zulueta, 1910
  • Lamippe pusilla Zulueta, 1908 (syn. Enalcyonium pusillum (Zulueta, 1908)
  • Lamippe rubicunda (Olsson, 1869) (syn. Enalcyonium rubicundum Olsson, 1869
  • Lamippe rubra Bruzelius, 1858
  • Lamippe setigera Zulueta, 1908 (syn. Enalcyonium setigerum (Zulueta, 1908)
  • Lamippe sympodii Zulueta, 1910 (syn. Enalcyonium sympodii (Zulueta, 1910)

    LAMIPPELLA Bouligand & Delamare Deboutteville, 1959

  • Lamippella acanellae Grygier, 1983
  • Lamippella delamarei Bouligand, 1965
  • Lamippella faurei Bouligand & Delamare Deboutteville, 1959

    LAMIPPINA Bouligand, 1960 (syn. Lamippulina Zulueta, 1908)

  • Lamippina aciculifera (Zulueta, 1908)
  • Lamippina aequalis Stock, 1973
  • Lamippina laubieri Bouligand, 1960

    LAMIPPULA Bouligand, 1966

  • Lamippula chattoni (Zulueta, 1908)
  • Lamippula duthiersi (Joliet, 1882)
  • Lamippula pallida (Zulueta, 1908)
  • Lamippula parva (Zulueta, 1908)

    LINARESIA Zulueta, 1908

  • Linaresia bouligandi Stock, 1979
  • Linaresia magna Grygier, 1980
  • Linaresia mammillifera Zulueta, 1908

    MAGNIPPE Stock, 1978

  • Magnippe caputmedusae Stock, 1978

    PTILOSARCOMA Williams, Anchaluisa, Boyko & McDaniel, 2018

  • Ptilosarcoma athyrmata Williams, Anchaluisa, Boyko & McDaniel, 2018

    SPHAERIPPE Grygier, 1980

  • Sphaerippe caligicola Grygier, 1980

    Alcyonicola Scott T. & Scott A., 1895 syn. Enalcyonium Olsson, 1869; Lamippulina Zulueta, 1908 syn. Lamippina Bouligand, 1960; Platypsyllus Scott T., 1902 syn- Jeanella Scott T., 1904.



    LEANIRICOLIDAE Huys, 2016

    LEANIRICOLA McIntosh, 1885

  • Leaniricola rotundata M'Intosh, 1885



    LUBBOCKIIDAE Huys & Böttger-Schnack, 1997

    Atrophia Huys & Böttger-Schnack, 1997

  • Atrophia glacialis (Sars G.O., 1900)
  • Atrophia minuta (Wolfenden, 1905)

    HAPLOPODIA Huys & Böttger-Schnack, 1997

  • Haplopodia petersoni (Heron & Damkaer, 1978)

    HOMEOGNATHIA Huys & Böttger-Schnack, 1997

  • Homeognathia brevis (Farran, 1908)
  • Homeognathia flemingi (Heron & Damkaer, 1978)

    LAITMATOBIUS Humes, 1987

  • Laitmatobius crinitus Humes, 1987

    LUBBOCKIA Claus, 1863

  • Lubbockia aculeata Giesbrecht, 1891
  • Lubbockia brevis Farran, 1908 (syn. Homeognathia brevis (Farran, 1908)
  • Lubbockia carinata Heron & Damkaer, 1978 (syn. Rhamphochela carinata (Heron & Damkaer, 1978)
  • Lubbockia extenuata Boxshall, 1977
  • Lubbockia flemingi Heron & Damkaer, 1978 (syn. Homeognathia flemingi (Heron & Damkaer, 1978)
  • Lubbockia forcipula Heron & Damkaer, 1978 (syn. Rhamphochela forcipula (Heron & Damkaer, 1978)
  • Lubbockia glacialis Sars G.O., 1900 (syn. Atrophia glacialis (Sars G.O., 1900)
  • Lubbockia marukawai Mori, 1937
  • Lubbockia minuta Wolfenden, 1905 (syn. Atrophia minuta (Wolfenden, 1905)
  • Lubbockia squillimana Claus, 1865
  • Lubbockia petersoni Heron & Damkaer, 1978 (syn. Haplopodia petersoni (Heron & Damkaer, 1978)
  • Lubbockia squillimana Claus, 1863 (?)
  • Lubbockia wilsonae Heron & Damkaer, 1969

    PSEUDOLUBBOCKIA Sars G.O., 1909

  • Pseudolubbockia dilatata Sars G.O., 1909

    RHAMPHOCHELA Huys & Böttger-Schnack, 1997

  • Rhamphochela carinata (Heron & Damkaer, 1978
  • Rhamphochela forcipula (Heron & Damkaer, 1978)



    MAKROSTROTIDAE Huys, Fatih, Ohtsuka & Llewellyn-Hughes, 2012

    MAKROSTROTOS Ho & Lin, 2006

  • Makrostrotos acuminatus Ho & Lin, 2006
  • Makrostrotos hamus Ho & Lin, 2006



    MESOGLICOLIDAE Zulueta, 1911

    MESOGLICOLA Quidor, 1906

  • Mesoglicola delagei Quidor, 1906



    MICRALLECTIDAE Huys, 2001

    MICRALLECTO Stock, 1971 (syn. Nannallecto Stock, 1973)

  • Micrallecto fusii (Stock, 1973)
  • Micrallecto uncinata Stock, 1971


    MEGALLECTO Gotto 1986 (?)

  • Megallecto thirioti Gotto 1986 (?)


    MYICOLIDAE Yamaguti, 1936

    PUSANOMYICOLA Lee et al., 2022

  • Pusanomyicola sensitivus Lee et al., 2022 [Korea]

    CONCHOCHERES Sars G.O., 1918 (syn. Panaietis malleolata (Sars G.O., 1918)

  • Conchocheres malleolatus Sars G.O., 1918 accepted as Panaietis malleolata (Sars G.O., 1918)

    CRUCISOMA Kabata, 1981

  • Crucisoma bernardi Kabata, 1981

    EXOSTRINCOLA Ho & Kim I.H., 1992

  • Exostrincola simplex (Humes, 1959)

    MYICOLA Wright R., 1885

  • Myicola clavator (Humes, 1959
  • Myicola formosanus Lin & Ho, 1999
  • Myicola gamoensis Kim I.H. & Sato, 2010
  • Myicola intumidus Kim I.H., 1997
    Myicola major (Williams, 1907) (syn. Leptinogaster major (Williams, 1907)
  • Myicola metisiensis Wright R., 1885
  • Myicola ostreae Hoshina & Sugiura, 1953
  • Myicola spinosa Pearse, 1947 (syn. Leptinogaster major (Williams, 1907)
  • Myicola tageli Pearse, 1947 (syn. Pseudomyicola spinosus (Raffaele & Monticelli, 1885); Pseudomyicola spinosus spinosus (Raffaele & Monticelli, 1885)
  • Myicola warneri Anderson, 1998

    OSTRINCOLA Wilson C.B., 1944

  • Ostrincola binoviger Kim I.H., 2009
  • Ostrincola breviseti Ho & Kim I.H., 1990
  • Ostrincola clavator Humes, 1959 (syn. Myicola clavator (Humes, 1959)
  • Ostrincola falcatus Humes, 1984
  • Ostrincola gracilis Wilson C.B., 1944
  • Ostrincola humesi Ho & Yoosukh, 1994
  • Ostrincola japonica Tanaka, 1961
  • Ostrincola koe Tanaka, 1961
  • Ostrincola patagonianus Humes, 1988
  • Ostrincola portonoviensis Reddiah, 1962
  • Ostrincola similis Lin & Ho, 1999
  • Ostrincola simplex Humes, 1959 (syn. Exostrincola simplex (Humes, 1959)

    PAROSTRINCOLA Humes & Boxshall, 1988

  • Parostrincola lingulae Humes & Boxshall, 1988

    PENGNA Ho & Kim I.H., 1992

  • Pengna bicornuta Ho & Kim I.H., 1992

    PSEUDOMYICOLA Yamaguti, 1936 (syn. Midicola Sowinsky, 1884 )

  • Pseudomyicola spinosus spinosus (Raffaele & Monticelli, 1885)



    MYTILICOLIDAE Bocquet & Stock, 1957

    CERASTOCHERES Monod & Dollfus, 1932

  • Cerastocheres trochicola Monod & Dollfus, 1932

    MYTICOLA Steuer, 1902 (syn. Noetiphilus Pearse, 1947; Piratasta Leigh-Sharpe, 1934

  • Mytilicola brachidontis (Yamaguti, 1939)
  • Mytilicola fimbriatus Humes & Ho, 1970
  • Mytilicola intestinalis Steuer, 1902
  • Mytilicola mactrae Hoshina & Kuwabara, 1959
  • Mytilicola orientalis Mori, 1935
  • Mytilicola ostrea Hoshina & Sugiura, 1953 (syn. Myicola ostreae Hoshina & Sugiura, 1953
  • Mytilicola porrecta Humes, 1954
  • Mytilicola virgatula (Leigh-Sharpe, 1934)

    PECTENOPHILUS Nagasawa, Bresciani & Lutzen, 1988

  • Pectenophilus ornatus Nagasawa, Bresciani & Lutzen, 1988 (Copepoda incertae sedis according to Huys et al., 2006)

    TROCHICOLA Dollfus, 1914

  • Trochicola enterica Dollfus, 1914
  • Trochicola japonica (japonicus?) Shimura & Kuwabara, 1984
  • Trochicola pectinidarum Tuzet & Ormières, 1962
  • Trochicola vermiformis Stock, 1959


    Noetiphilus Pearse, 1947 syn. Mytilicola Steuer, 1902; Piratasta Leigh-Sharpe, 1934 syn. Mytilicola Steuer, 1902



    NEREICOLIDAE Claus, 1875

    ANOMOPSYLLUS Sars G.O., 1921

  • Anomopsyllus abyssorum Laubier, 1988
  • Anomopsyllus bifurcus Kim I.H., Sikorski, O'Reilly & Boxshall, 2013
  • Anomopsyllus geminus Kim I.H., Sikorski, O'Reilly & Boxshall, 2013
  • Anomopsyllus hamiltonae Kim I.H., Sikorski, O'Reilly & Boxshall, 2013
  • Anomopsyllus pranizoides Sars G.O., 1921

    NEREICOLA Keferstein, 1863 (syn. Nereidicola Grube, 1869)

  • Nereicola concinna Scott T., 1902 (syn. Sigecheres concinna (Scott T., 1902)
  • Nereicola ovata Keferstein, 1863

    PHERMA Wilson C.B., 1923 (syn. Pestifer Wilson C.B., 1944 )

  • Pherma curticaudatum Wilson C.B., 1923

    SELIOIDES Levinsen, 1878 (syn. Cancerina Scott T., 1901)

  • Selioides bocqueti Carton, 1963
  • Selioides bolbroei Levinsen, 1878
  • Selioides bulbifer Stock, 1986
  • Selioides capensis Stock, 1996
  • Selioides confusus (Scott T., 1901) (syn. Selioides bolbroei Levinsen, 1878)
  • Selioides guineensis Carton & Laubier, 1974
  • Selioides tardus Gravier, 1912

    SELIUS Krøyer, 1837

  • Selius bilobus Krøyer, 1837

    SIGECHERES Bresciani, 1964

  • Sigecheres concinna (Scott T., 1902)

    VECTORIELLA Stock, 1968

  • Vectoriella gabesensis Kim I.H., Sikorski, O'Reilly & Boxshall, 2013
  • Vectoriella marinovi Stock, 1968
  • Vectoriella ramosae Laubier & Carton, 1973

    Cancerina Scott T., 1901 syn. Selioides Levinsen, 1878; Chelonidium Claus, 1875 syn. Chelonidiformis Hesse, 1869; Nereidicola Grube, 1869 syn.Nereicola Keferstein, 1863; Pestifer Wilson C.B., 1944 syn. Pherma Wilson C.B., 1923.



    OCTOPICOLIDAE Humes & Boxshall, 1996

    OCTOPICOLA Humes, 1957

    "The genus Octopicola Humes, 1957 was included in the family Lichomolgidae Kossmann, 1877 in some earlier publications (Humes & Stock, 1972, 1973). Because it differs from the other members of the Lichomolgidae by retaining the primitive 6-segmented female urosome, Humes & Boxshall (1996) established a new family, Octopicolidae Humes & Boxshall, 1996, to accommodate it"

  • Octopicola huanghaiensis Du, Dong & Sun, 2018
  • Octopicola regalis Humes, 1974
  • Octopicola stocki Humes, 1963
  • Octopicola superba superba Humes, 1957
  • Octopicola superbus antillensis Stock et al. (?)



    ONCAEIDAE Giesbrecht, 1893

    ARCHIONCAEA Böttger-Schnack & Huys, 1997

  • Archioncaea arabica Böttger-Schnack & Huys, 1997

    CONAEA Giesbrecht, 1891

  • Conaea atlantica (Shmeleva, 1967) (syn. Oncaea atlantica Shmeleva, 1967
  • Conaea expressa (Gordeeva K.T., 1973)
  • Conaea gracilis (Dana, 1853) (syn, Oncaea gracilis (Dana, 1852)
  • Conaea hispida Heron, 1977
  • Conaea rapax Giesbrecht, 1891
  • Conaea succurva Heron, 1977

    EPICALYMMA Heron, 1977

  • Epicalymma ancora (Gordeeva K.T., 1973)
  • Epicalymma brittoni Heron, English & Damkaer, 1984
  • Epicalymma bulbosa Böttger-Schnack, 2009
  • Epicalymma exigua (Farran, 1908)
  • Epicalymma schmitti Heron, 1977
  • Epicalymma umbonata Heron, 1977
  • Epicalymma vervoorti Heron, English & Damkaer, 1984

    MONOTHULA Böttger-Schnack & Huys, 2001

  • Monothula subtilis (Giesbrecht, 1893)


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