SPLANCHNOTROPHIDAE Norman & Scott T., 1906
Splanchnotrophidae are highly modified
copepods which exclusively utilize a variety
of marine opisthobranch gastropods as hosts.
ARTHURIUS Huys, 2001
Arthurius bunakenensis Salmen, Kaligis, Mamangkey & Schrödl, 2008
Arthurius elysiae (Jensen, 1990)
Arthurius gibbosa Anton, Schories, Jörger, Kaligis & Schrödl, 2016
CERATOSOMICOLA Huys, 2001
Ceratosomicola coia Salmen, Wilson & Schrödl, 2008
Ceratosomicola delicata Salmen, Wilson & Schrödl, 2008
Ceratosomicola japonica Uyeno & Nagasawa, 2012
Ceratosomicola mammilata Salmen, Wilson & Schrödl, 2008
Ceratosomicola oki Uyeno et al., 2021 [Japan]
Ceratosomicola sacculata (O'Donoghue, 1924)
ISMAILA Bergh, 1868
Ismaila aliena Haumayr & Schrödl, 2003
Ismaila androphila Haumayr & Schrödl, 2003
Ismaila belciki Ho, 1987
Ismaila chaihuiensis Anton, Schories, Jörger, Kaligis & Schrödl, 2016
Ismaila damnosa Haumayr & Schrödl, 2003
Ismaila genalis Anton, Schories, Jörger, Kaligis & Schrödl, 2016
Ismaila jenseniana Haumayr & Schrödl, 2003
Ismaila magellanica Haumayr & Schrödl, 2003
Ismaila monstrosa Bergh, 1867
Ismaila obtusa Haumayr & Schrödl, 2003
Ismaila occulta Ho, 1981
Ismaila robusta Haumayr & Schrödl, 2003
Ismaila socialis Haumayr & Schrödl, 2003
Ismaila volatilis Anton, Schories, Jörger, Kaligis & Schrödl, 2016
LOMANOTICOLA Scott T. & Scott A., 1895
Lomanoticola brevipes (Hancock & Norman, 1863)
Lomanoticola insolens Scott T. & Scott A., 1895
Lomanoticola nishiharai Uyeno & Hirose, 2018
MAJIMUN Uyeno & Nagasawa, 2012
Majimun shirakawai Uyeno & Nagasawa, 2012
SPLANCHNOTROPHUS Hancock & Norman, 1863
Splanchnotrophus angulatus Hecht, 1893
Splanchnotrophus brevipes Hancock & Norman, 1863 (syn. Lomanoticola brevipes (Hancock & Norman, 1863)
Splanchnotrophus dellachiajei Delamare Deboutteville, 1951
Splanchnotrophus elysiae Jensen, 1990 (syn. Arthurius elysiae (Jensen, 1990)
Splanchnotrophus gracilis Hancock & Norman, 1863
Splanchnotrophus helianthus Uyeno & Nagasawa, 2012
Splanchnotrophus imagawai Uyeno & Nagasawa, 2012
Splanchnotrophus insolens (Scott, T. & Scott A., 1895) (syn. Lomanoticola insolens Scott T. & Scott A., 1895)
Splanchnotrophus sacculatus O'Donoghue, 1924 (syn. Ceratosomicola sacculata (O'Donoghue, 1924)
Splanchnotrophus willemi Canu, 1891
CHONDRACANTHIDAE Milne Edwards, 1840 [syn. Andreinidae Yamaguti, 1963;
Lernaeosoleidae Hogans & Benz, 1990; Pharodidae Illg, 1948), Lichomolgoidea (Cyclopoida) according to Huys et al., 2006]
"The family Chondracanthidae H. Milne Edwards, 1840 is composed of
192 species under 50 valid genera (Tang et al., 2007; Braicovich
et al., 2013; Gomez et al., 2018; Ohtsuka et al., 2019; Walter and Boxshall,
2019). The genus Acanthochondria Oakley, 1930 is the largest genus in this
family with 52 valid species, followed by Chondracanthus Delaroche, 1811
represented by 41 valid species (Tang et al., 2007; Braicovich et al., 2013;
Gomez et al., 2018; Ohtsuka et al., 2019)" (from:
ACANTHOCANTHOPSIS Heegaard, 1945
Acanthocanthopsis chilomycteri (Thomson, 1889)
Acanthocanthopsis quadrata Heegaard, 1945
ACANTHOCHONDRIA Oakley, 1930 (syn. Entomoda Lamark, 1818; Pterochondria Ho, 1973; Rylovia Markevich, 1940) Acanthochondria Oakley, 1927 is the largest genus of the family Chondracanthidae Edwards, 1840, containing 48 valid
species (Alves et al. 2003, Kalman 2003, Østergaard 2003,
Braicovich and Timi 2009)
Acanthochondria alatalongicollis Heegaard, 1940
Acanthochondria alleni Tang, Kalman & Ho, 2010
Acanthochondria albigutta Pearse, 1952 [Australia] (?)
Acanthochondria bicornis Shiino, 1955
Acanthochondria brevicorpa Yü, 1935
Acanthochondria chilomycteri (Thomson G.M., 1890) (syn. Acanthocanthopsis chilomycteri (Thomson, 1889)
Acanthochondria clavata (Bassett-Smith, 1896)
Acanthochondria constricta Shiino, 1955
Acanthochondria cornuta (Müller O.F., 1776)
Acanthochondria crassicornis (Krøyer, 1837) (syn. Heterochondria crassicornis (Krøyer, 1837)
Acanthochondria krishnai Aneesh et al., 2020
Acanthochondria cyclopsetta Pearse, 1952
Acanthochondria cynoglottidis (Thompson I.C. & Scott A., 1903) (syn. Heterochondria cynoglottidis (Thompson I.C. & Scott A., 1903)
Acanthochondria deltoidea (Fraser, 1920) (syn. Chondracanthus deltoideus Fraser, 1920 )
Acanthochondria diastema Kabata, 1965
Acanthochondria dilatata Shiino, 1955
Acanthochondria dojirii Kabata, 1984
Acanthochondria elongata (Bassett-Smith, 1898)
Acanthochondria epachthes (Wilson C.B., 1908)
Acanthochondria eptratreti Cheng, Luo, Dai & Shih, 2014
Acanthochondria exilipes Wilson C.B., 1932
Acanthochondria fissicauda Shiino, 1955
Acanthochondria fraseri Ho, 1972
Acanthochondria galerita (Rathbun, 1886)
Acanthochondria glandiceps Shiino, 1955
Acanthochondria helicoleni Cantatore & Timi, 2010
Acanthochondria hippoglossi Kabata, 1987
Acanthochondria hoi Kalman, 2003
Acanthochondria incisa Shiino, 1955
Acanthochondria inimici Yamaguti, 1939
Acanthochondria kajika Ho & Kim I.H., 1996
Acanthochondria krishnai Aneesh, Helna & Kumar, 2020
Acanthochondria laemonemae Capart, 1959
Acanthochondria lepidionis Barnard, 1955
Acanthochondria lilianae Cantatore, Lanfranchi & Timi, 2011
Acanthochondria limandae (Krøyer, 1863)
Acanthochondria longicephalus Yü & Wu, 1932 (syn. Heterochondria longicephala (Yü & Wu, 1932)
Acanthochondria longifrons Shiino, 1955
Acanthochondria macrocephala Gusev, 1951
Acanthochondria margolisi Kabata, 1984
Acanthochondria ophidii (Krøyer, 1863)
Acanthochondria oralis Yamaguti, 1939
Acanthochondria palpifera (Wilson C.B., 1912) (syn. Chondracanthus palpifer Wilson C.B., 1912)
Acanthochondria phycidis (Rathbun, 1886)
Acanthochondria pingi (Yü & Wu, 1932)
Acanthochondria platycephali Heegaard, 1940
Acanthochondria priacanthi Shiino, 1964
Acanthochondria psetti (Krøyer, 1863) (syn. Chondracanthus psetti Krøyer, 1863)
Acanthochondria radiata (Lamarck, 1818) (syn. Chondracanthodes radiatus (Müller O.F., 1776)
Acanthochondria rectangularis (Fraser, 1920)
Acanthochondria sagitta Alarcos & Timi, 2011
Acanthochondria salmonea (Lamarck, 1818) (syn. Salmincola salmonea (Linnaeus, 1758)
Acanthochondria serrani Braicovich & Timi, 2009
Acanthochondria shawi Yü, 1935
Acanthochondria sicyasis (Krøyer, 1863)
Acanthochondria sixteni (Wilson C.B., 1922)
Acanthochondria soleae (Krøyer, 1838)
Acanthochondria solida Gusev, 1951 (syn. Chondracanthus solidus (Gusev, 1951)
Acanthochondria spirigera Shiino, 1955
Acanthochondria tasmaniae Heegaard, 1962
Acanthochondria tchangi Yü, 1935
Acanthochondria tenuis Pearse, 1952 (syn. Berea ancoralis (Bere, 1936) (syn. Bereacanthus ancoralis (Bere, 1936)
Acanthochondria triangularis Alves, Luque & Paraguassu, 2003
Acanthochondria triglae Herrera-Cubilla & Raibaut, 1990
Acanthochondria uranoscopi Ho & Kim I.H., 1995
Acanthochondria vancouverensis Kabata, 1984
Acanthochondria yui Shiino, 1964
Acanthochondria zebriae Ho, Kim I.H. & Kumar, 2000 accepted as Heterochondria zebriae (Ho, Kim I.H. & Kumar, 2000)
KEY TO SPECIES OF ACANTHOCONDRIA
AVATAR Aneesh et al, 2024
Avatar mirabilis Aneesh et al, 2024 [Japan] (?)
Avatar nishidai Aneesh et al, 2024 [Japan]
KOKESHIOIDES Aneesh et al, 2024
Kokeshioides surugaensis Aneesh et al, 2024 [Japan]
ACANTHOCHONDRITES Oakley, 1930
Acanthochondrites annulatus (Olsson, 1869)
ANDREINA Brian, 1939
Andreina synapturae Brian, 1939
APODOCHONDRIA Ho & Dojiri, 1988
Apodochondria medusae Ho & Dojiri, 1988
ARGENTINOCHONDRIA Etchegoin, Timi & Sardella, 2003
Argentinochondria patagonensis Etchegoin, Timi & Sardella, 2003
AUCHENOCHONDRIA Dojiri & Perkins, 1979
Auchenochondria lobosa Dojiri & Perkins, 1979
BACTROCHONDRIA Ho, Kim I.H. & Kumar, 2000
Bactrochondria formosana Ho, Lin & Liu, 2011
Bactrochondria hoi (Pillai, 1985)
Bactrochondria longitruncus (Yamaguti, 1939)
Bactrochondria papilla Ho, Kim I.H. & Kumar, 2000
Bactrochondria tuase Ho, Lin & Liu, 2011
BEREACANTHUS Huys, 2009 (syn. Berea Yamaguti, 1963)
Bereacanthus ancoralis (Bere, 1936)
Bereacanthus clava (Ho & Sey, 1997)(syn. Berea clava Marsh 1944)
BLIAS Krøyer, 1863 (syn. Trichthacerus Krøyer, 1863; Triphyllacanthus Oakley, 1930; Tuccopsis Pearse, 1952)
Blias ancoralis (Bere, 1936) (syn. Bereacanthus ancoralis (Bere, 1936)
Blias marplatensis Timi, Etchegoin & Lanfranchi, 2004
Blias prionoti Krøyer, 1863
BOBKABATA Hogans & Benz, 1990
Bobkabata kabatabobbus Hogans & Benz, 1990
BRACHIOCHONDRIA Shiino, 1957
Brachiochondria higanfugu Yamaguti & Yamasu, 1959
Brachiochondria murtii Rangnekar, 1977
Brachiochondria pinguis Shiino, 1957
BRACHIOCHONDRITES Markevich, 1940
Brachiochondrites longicollis Markevich, 1940
BRASILOCHONDRIA Thatcher & Pereira Júnior, 2004
Brasilochondria riograndensis Thatcher & Pereira Júnior, 2004
CERATOCHONDRIA Yu, 1935
Ceratochondria brevicollis (Krøyer, 1863)
Ceratochondria hoi Pillai, 1985 (syn. Bactrochondria hoi (Pillai, 1985)
Ceratochondria longitruncus (Yamaguti, 1939) (syn. Bactrochondria longitruncus (Yamaguti, 1939)
CHELONICHONDRIA Ho, 1994
Chelonichondria okamurai Ho, 1994
Subfamily CHONDRACANTHINAE Leigh-Sharpe & Oakley, 1927
CHONDRACANTHODES Wilson C.B., 1932
Chondracanthodes bulbosus Kabata, 1965
Chondracanthodes deflexus Wilson C.B., 1932
Chondracanthodes lotellae (Thomson G.M., 1890) ( syn. Chondracanthus irregularis Fraser, 1920)
Chondracanthodes radiatus (Müller O.F., 1776)
Chondracanthodes tuberofurcatus Kabata & Gusev, 1966
CHONDRACANTHUS Delaroche, 1811 (syn. Barnardia Yamaguti, 1963; Chondracanthopsis Oakley, 1930; Disphaerocephalus Oakley, 1930; Lernacantha Blainville, 1822;
Protochondracanthoides Yamaguti, 1963)
Chondracanthus alatus Heller, 1865 (syn. Protochondracanthus alatus (Heller, 1865)
Chondracanthus angustatus Heller, 1865
Chondracanthus annulatus Olsson, 1869 (syn. Acanthochondrites annulatus (Olsson, 1869)
Chondracanthus australis Ho, 1991
Chondracanthus barnardi Ho, 1972
Chondracanthus brevicollis Krøyer, 1863 (syn. Ceratochondria brevicollis (Krøyer, 1863)
Chondracanthus briani Yü & Wu, 1932 (syn. Acanthochondria briani (Yü & Wu, 1932)
Chondracanthus brotulae Capart, 1959
Chondracanthus bulbosus (Kabata, 1965) (syn. Chondracanthodes bulbosus Kabata, 1965
Chondracanthus chilensis Wilson C.B., 1918 (syn. Acanthochondria phycidis (Rathbun, 1886)
Chondracanthus chilomycteri Thomson G.M., 1890 (syn. Acanthocanthopsis chilomycteri (Thomson, 1889)
Chondracanthus cynoglottidis Thompson I.C. & Scott A., 1903 (syn. Heterochondria cynoglottidis (Thompson I.C. & Scott A., 1903)
Chondracanthus clavatus Bassett-Smith, 1896 (syn. Acanthochondria clavata (Bassett-Smith, 1896)
Chondracanthus colligens Barnard, 1955
Chondracanthus cornutus (Müller O.F., 1776) (syn. Acanthochondria cornuta (Müller O.F., 1776)
Chondracanthus cottunculi Rathbun, 1886
Chondracanthus crassicornis Krøyer, 1837 (syn. Heterochondria crassicornis (Krøyer, 1837)
Chondracanthus deltoideus Fraser, 1920
Chondracanthus depressus Scott T., 1905 (syn. Acanthochondria cornuta (Müller O.F., 1776)
Chondracanthus dibranchi Go´mez, Aguirre-Villaseñor & Morales-Serna, 2018
Chondracanthus elongatus Bassett-Smith, 1898 (syn. Acanthochondria elongata (Bassett-Smith, 1898)
Chondracanthus epachthes Wilson C.B., 1908 (syn. Acanthochondria epachthes (Wilson C.B., 1908)
Chondracanthus flurae Krøyer, 1863 (syn, Acanthochondria cornuta (Müller O.F., 1776)
Chondracanthus galeritus Rathbun, 1886 (syn. Acanthochondria galerita (Rathbun, 1886)
Chondracanthus gobinus (Müller O.F., 1776) (syn. Diocus gobinus (Müller O.F., 1776)
Chondracanthus goldsmidi Tang, Andrews & Cobcroft, 2007
Chondracanthus gracilis Fraser, 1920
Chondracanthus grandigenitalis Yü & Wu, 1932 (syn. Acanthochondria grandigenitalis (Yü & Wu, 1932)
Chondracanthus gurnardi Olsson, 1869 (syn. Acanthochondria gurnardi (Olsson, 1869)
Chondracanthus heterostichi Ho, 1972
Chondracanthus hoi Braicovich, Lanfranchi, Incorvaia, Inés & Timi, 2013
Chondracanthus horridus Heller, 1865
Chondracanthus inflatus Bainbridge, 1909 (syn. Acanthochondrites annulatus (Olsson, 1869)
Chondracanthus irregularis Fraser, 1920
Chondracanthus janebennettae Causey, 1953
Chondracanthus kabatai Aneesh, Helna & Kumar, 2020
Chondracanthus laevirajae Della Valle, 1881 (syn. Acanthochondrites annulatus (Olsson, 1869)
Chondracanthus lepidionis Kabata, 1970
Chondracanthus lepophidii Ho, 1974
Chondracanthus limandae Krøyer, 1863 (syn. Acanthochondria limandae (Krøyer, 1863)
Chondracanthus longicephalus Yü & Wu, 1932 (syn. Heterochondria longicephala (Yü & Wu, 1932)
Chondracanthus lophii Johnston, 1836
Chondracanthus macrurus Brady, 1883 (syn. Lateracanthus macrurus (Brady, 1883)
Chondracanthus merluccii (Holten, 1802)
Chondracanthus multituberculatus (Markevich, 1956)
Chondracanthus narium Kabata, 1969
Chondracanthus neali Leigh-Sharpe, 1930
Chondracanthus ninnii Richiardi, 1882 (syn. Pharodes ninnii (Richiardi, 1882)
Chondracanthus nodosus (Müller O.F., 1776)
Chondracanthus ophidii Krøyer, 1863 (syn. Acanthochondria ophidii (Krøyer, 1863)
Chondracanthus ornatus Scott T., 1900
Chondracanthus pallidus Beneden, 1870 (syn. Acanthochondrites annulatus (Olsson, 1869)
Chondracanthus palpifer Wilson C.B., 1912
Chondracanthus parvus Ho et al., 2005 (?)
Chondracanthus phycidis Rathbun, 1886 (syn. Acanthochondria phycidis (Rathbun, 1886)
Chondracanthus pingi Yü & Wu, 1932 (syn. Acanthochondria pingi (Yü & Wu, 1932)
Chondracanthus pinguis Wilson C.B., 1912
Chondracanthus polymixiae Yamaguti, 1939
Chondracanthus psetti Krøyer, 1863
Chondracanthus pusillus Kabata, 1968
Chondracanthus quadratus (Heegaard, 1945) (syn. Acanthocanthopsis quadrata Heegaard, 1945)
Chondracanthus radiatus (Müller O.F., 1776) (syn. Chondracanthodes radiatus (Müller O.F., 1776)
Chondracanthus rectangularis Fraser, 1920 (syn. Acanthochondria rectangularis (Fraser, 1920)
Chondracanthus rickettsi (Wilson C.B., 1935) (syn. Chondracanthodes deflexus Wilson C.B., 1932
Chondracanthus shiinoi Yamaguti, 1963
Chondracanthus sicyasis Krøyer, 1863 (syn. Acanthochondria sicyasis (Krøyer, 1863)
Chondracanthus similis Yü & Wu, 1932 (syn. Heterochondria similis (Yü & Wu, 1932)
Chondracanthus sixteni Wilson C.B., 1922 (syn. Acanthochondria sixteni (Wilson C.B., 1922)
Chondracanthus soleae Krøyer, 1838 (syn. Acanthochondria soleae (Krøyer, 1838)
Chondracanthus solidus (Gusev, 1951)
Chondracanthus theragrae Yamaguti, 1939
Chondracanthus triglae Guérin-Méneville, 1837 (syn. Lernentoma asellina (Linnaeus, 1758)
Chondracanthus trilobatus Pillai, 1964 (syn. Protochondracanthus trilobatus (Pillai, 1964)
Chondracanthus triventricosus Sekerak, 1970
Chondracanthus tuberculatus Nordmann, 1832
Chondracanthus wilsoni Ho, 1971
Chondracanthus yabei Ho, J.S., Kim I.H. & Nagasawa, 2005
Chondracanthus yanezi Atria, 1980
Chondracanthus zei Delaroche, 1811
CRYPTOCHONDRIA Izawa, 1971
Cryptochondria tricaudata Izawa, 1971
DIOCUS Krøyer, 1863 (syn. Parapharodes Shiino, 1960)
Diocus clini Vaney & Conte, 1900 (syn. Pharodes clini (Vaney & Conte, 1900)
Diocus frigidus Hansen, 1923
Diocus gobinus (Müller O.F., 1776)
Diocus lycenchelus Hogans & Sulak, 1992
Diocus sadoensis (Shiino, 1960)
Diocus semilunaris (Kabata & Gusev, 1966)
HETEROCHONDRIA Yu, 1935
Heterochondria atypica Ho, 1972
Heterochondria crassicornis (Krøyer, 1837)
Heterochondria cynoglottidis (Thompson I.C. & Scott A., 1903)
Heterochondria longa Tripathi, 1960
Heterochondria longicephala (Yü & Wu, 1932)
Heterochondria longitruncus (Yamaguti, 1939) (syn. Bactrochondria longitruncus (Yamaguti, 1939)
Heterochondria orientalis Moon & Soh, 2013
Heterochondria petila Ho, Kim I.H. & Kumar, 2000
Heterochondria pillaii Ho, 1970
Heterochondria similis (Yü & Wu, 1932)
Heterochondria zebriae (Ho, Kim I.H. & Kumar, 2000)
HOIA Avdeev & Kazachenko, 1986
Hoia hoi Avdeev & Kazachenko, 1985
HUMPHREYSIA Leigh-Sharpe, 1934
Humphreysia floreata Leigh-Sharpe, 1934
Humphreysia hoi Do & Kasahara, 1982 (syn. Prochondracanthopsis quadricornutus Shiino, 1960
IMMANTHE Leigh-Sharpe, 1934
Immanthe campanulata Leigh-Sharpe, 1934
JUANETTIA Wilson C.B., 1921
Juanettia continentalis Villalba & Fernandez, 1985
Juanettia cornifera Wilson C.B., 1921
JUSHEYHOEA Villalba & Fernandez, 1985
Jusheyhoea macrura Villalba & Fernandez, 1985
Jusheyhoea moseri Kabata, 1991
Jusheyhoea ryukyuensis Ho, 1994
LAGOCHONDRIA Ho & Dojiri, 1988
Lagochondria nana Ho & Dojiri, 1988
Lagochondria quadrata Ohtsuka & Nitta, in Ohtsuka, Nitta, Nakamura & Maekawa, 2019
LATERACANTHUS Kabata & Gusev, 1966
Lateracanthus curtus Kabata, 1993
Lateracanthus novus Kabata, 1992
Lateracanthus quadripedis Kabata & Gusev, 1966
LERNAEOSOLEA Wilson C.B., 1944
Lernaeosolea lycodis Wilson C.B., 1944
LERNENTOMA Blainville, 1822 (syn. Lernaeomyzon Blainville, 1822 )
Lernentoma asellina (Linnaeus, 1758)
Lernentoma cornuta (Müller O.F., 1776) (syn. Chondracanthus cornutus (Müller O.F., 1776)
Lernentoma dufresnii Blainville, 1822 (syn. Chondracanthus lophii Johnston, 1836)
Lernentoma gobina (Müller O.F., 1776) (syn. Chondracanthus gobinus (Müller O.F., 1776)
Subfamily LEMENTOMINAE Leigh-Sharpe & Oakley, 1927
MARKEVITCHIELINUS Titar, 1975
Markevitchielinus anchoratus Titar, 1975
MECADEROCHONDRIA Ho & Dojiri, 1987
Mecaderochondria pilgrimi Ho & Dojiri, 1987
MEDESICASTE Krøyer, 1863
Medesicaste asellinum (Linnaeus, 1758) (syn. Lernentoma asellina (Linnaeus, 1758)
Medesicaste penetrans Heller, 1865
Medesicaste triglarum Krøyer, 1863
NEOBRACHIOCHONDRIA Kabata, 1969
Neobrachiochondria quadrata Kabata, 1969
PARAPERCICOLA Ho, Liu & Lin, 2011
Parapercicola formosana Ho, Liu & Lin, 2011
Parapercicola inflata Ho, Liu & Lin, 2013
PHARODES Wilson C.B., 1935
Pharodes banyulensis Delamare Deboutteville & Nunes-Ruivo, 1952
Pharodes biakensis Illg, 1948
Pharodes clini (Vaney & Conte, 1900)
Pharodes ninnii (Richiardi, 1882)
Pharodes tortugensis Wilson C.B., 1935
PRAECIDOCHONDRIA Kabata, 1968
Praecidochondria galatheae Kabata, 1968
Praecidochondria setoensis Izawa, 1975
Key to subfamilies and genera
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