MIRACIIDAE Dana, 1846 (Willen, 2002) (syn. DIOSACCIDAE Sars, 1906)

DIOSACCINAE Sars,1906


DIOSACCUS Boeck, 1872 [syn. PARADIOSACCUS Lang, 1944]

  • Diosaccus abyssi Boeck, 1873
  • Diosaccus borborocoetus Jakobi, 1954 [Brazil; marine coastal waters]
  • Diosaccus dentatus (Thompson & A. Scott, 1903 [Sri Lanka; marine]
  • Diosaccus ezoensis Ito, 1974 [Japan; marine]
  • Diosaccus hamiltoni (Thompson & A. Scott, 1903) [Sri Lanka; marine]
  • Diosaccus koreanus Lim et al., 2020 [Korea; marine]
  • Diosaccus longirostris (Claus, 1863) accepted as Paramphiascopsis longirostris (Claus, 1863)
  • Diosaccus monardi Sewell, 1940 [India; marine] *
  • Diosaccus propinquus Scott T. & Scott A., 1893 accepted as Pseudodiosaccus propinquus (Scott T. & Scott A., 1893)
  • Diosaccus rebus (Sewell, 1940) [India?; ?]
  • Diosaccus robustus (Thompson & A. Scott, 1903 [Sri Lanka; marine]
  • Diosaccus ruber Brian, 1923
  • Diosaccus sordidus Lang, 1948
  • Diosaccus spinatus Campbell, 1929 [north America, ?]
  • Diosaccus tenuicornis (Claus, 1863) [Europe, north Africa, Americas, coastal marine waters, muddy sand]
  • Diosaccus truncatus Gurney, 1927 [Egypt; marine]
  • Diosaccus valens (Gurney, 1927) [Egypt; marine]
  • Diosaccus varicolor varicolor (Farran, 1913) [Europe; marine]
  • Diosaccus varicolor biarticulatus (Monard, 1924) [France, Italy, Algerie; marine]
  • Diosaccus varicolor pentasetosus (Noodt, 1955) [Marmara Sea]

    * synonym : Diosaccus monardi f. major Krishnaswamy (1957) (Wells & Rao, 1987)


    ANTIBOREODIOSACCUS Lang, 1944

  • Antiboreodiosaccus crassus (Giesbrecht, 1902) [Chile; marine]


    DACTYLOPODAMPHIASCOPSIS Lang, 1944

  • Dactylopodamphiascopsis latifolius (Sars, 1909) [north Europe, north America; marine]


    HELMUTKUNZIA Wells & Rao, 1976

  • Helmutkunzia hartmannorum (Kunz, 1971) [Angola; marine]
  • Helmutkunzia variabilis Wells & Rao, 1987 [Andaman Islands; marine]
  • Helmutkunzia xiameniensis Mu & Huys, 2021 [China]


    IALYSUS Brian, 1927 (syn. Jalysus Sewell, 1940)

  • Ialysus investigatoris Sewell, 1940 (syn. Parialysus investigatoris (Sewell, 1940)
  • Ialysus proximus Sewell, 1940 (syn.Parialysus proximus (Sewell, 1940)
  • Ialysus rufus Brian, 1927 [Egypt, Rhodes; marine]


    PARIALYSUS Nicholls, 1941

  • Parialysus proximus (Sewell, 1940)
  • Parialysus robustus (Nicholls, 1941) [Australia?; marine]
  • Parialysus investigatoris (Sewell, 1940)


    METAMPHIASCOPSIS Lang, 1944

  • Metamphiascopsis banyulensis (Monard, 1928) [France; marine]
  • Metamphiascopsis hirsutus hirsutus (Thompson & A. Scott, 1903) [Sri Lanka, Europe, Algerie; marine] (syn. Metamphiascopsis hirsutus hirsutus (Thompson I.C. & Scott A., 1903)
  • Metamphiascopsis hirsutus bermudae (Willey, 1930) [Bermuda; marine]
  • Metamphiascopsis nicobaricus (Sewell, 1940) [India?; marine]
  • Metamphiascopsis monardi (lang, 1934) [Campbell island; marine]


    MISCEGENUS Wells, Hicks & Coull, 1982

  • Miscegenus heretaunga Wells, Hicks & Coull, 1982 [New Zealand; marine coastal waters]


    ONYCHOSTENHELIA Ito, 1979

    "The genus Onychostenhelia was proposed by Itô (1979) for a new species, O. falcifera, based on five specimens dredged from a sandy substratum off Oshoro, Hokkaido, Japan. Its relationships clearly lie with the subfamily Stenheliinae (Miraciidae) because of the dorsolateral position of the female genital apertures on the genital double-somite.

    Itô (1979) suggested that, within this subfamily, Onychostenhelia most closely resembled Pseudostenhelia in the segmentation and armature of the swimming legs, and that the female P5 was similar to that found in Delavalia cornuta (Lang, 1936).

    However, the genus clearly differed from any other harpacticoid in the peculiar transformation of the P4 exopod in the male" [from: Huys & Mu, 2008].


  • Onychostenhelia falcifera Ito, 1979 [Japan; marine]
  • Onychostenhelia bispinosa Huys & Mu 2008 [China; marine]



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