"Monstrilloids are semi-parasitic copepods with a complex life cycle that includes an endoparasitic juvenile phase and a free-living planktonic adult phase. Early infective nauplii are also planktonic, but soon infect hosts. Juveniles are known to infect various marine invertebrates such as polychaetes, mollusks, and sponges (Boxshall and Halsey 2004; Huys et al. 2007; Suárez-Morales et al. 2014). Adults are generally scarce in open marine water, but are known to occur frequently in reef environments, especially at nighttime (Sale et al. 1976; Sekiguchi 1982; Suárez-Morales 2001; Grygier and Ohtsuka 2008).The order Monstrilloida Sars, 1901 currently contains a single family, Monstrillidae Dana, 1849, consisting of about 160 nominal species in six valid genera: Monstrilla Dana, 1849; Cymbasoma ? T ompson, 1888; Monstrillopsis Sars, 1921; Maemonstrilla Grygier & Ohtsuka, 2008; Australomonstrillopsis Suárez-Morales & McKinnon, 2016; and Caromiobenella Jeon, Lee & Soh, 2018" (from Jeon et al., 2019)


MONSTRILLIDAE Dana, 1849


AUSTRALOMONSTRILLOPSIS Suárez-Morales & McKinnon, 2014

  • Australomonstrillopsis crassicaudata Suárez-Morales & McKinnon, 2014

    CAROMIOBENELLA Jeon, W Lee & Soh, 2018

  • Caromiobenella arctica (Davis & Green, 1974)
  • Caromiobenella brasiliensis (Dias & Suárez-Morales, In: Suárez-Morales & Dias, 2000)
  • Caromiobenella castorea Jeon, W Lee & Soh, 2018
  • Caromiobenella hamatapex (Grygier & Ohtsuka, 1995)
  • Caromiobenella helgolandica (Claus, 1863)
  • Caromiobenella ohtsukai Jeon, W Lee & Soh, 2019
  • Caromiobenella patagonica (Suárez-Morales, Ramírez & Derisio, 2008)
  • Caromiobenella polluxea Jeon, W Lee & Soh, 2018
  • Caromiobenella pygmaea (Suárez-Morales, 2000)
  • Caromiobenella serricornis (Sars G.O., 1921)

    CYMBASOMA Thompson I.C., 1888

  • Cymbasoma agoense Sekiguchi, 1982
  • Cymbasoma alvaroi Suárez-Morales & Carrillo, 2013
  • Cymbasoma annulocolle Suárez-Morales & McKinnon, 2016
  • Cymbasoma apicale Suárez-Morales & McKinnon, 2016
  • Cymbasoma astrolabe Suárez-Morales & McKinnon, 2016
  • Cymbasoma bali Desai & Krishnaswamy, 1962
  • Cymbasoma belizense Suárez-Morales, Vasquez-Yeomans & Santoya, 2020
  • Cymbasoma bidentatum Suárez-Morales & McKinnon, 2016
  • Cymbasoma bitumidum Suárez-Morales & McKinnon, 2016
  • Cymbasoma boonwurrungorum Suárez-Morales & McKinnon, 2014
  • Cymbasoma bowmani Suárez-Morales & Gasca, 1998
  • Cymbasoma boxshalli (Suárez-Morales, 1993)
  • Cymbasoma buckleyi Suárez-Morales & McKinnon, 2016
  • Cymbasoma bullatum (Scott A., 1909)
  • Cymbasoma californiense Suárez-Morales & Palomares-Garcia, 1999
  • Cymbasoma chelemense Suárez-Morales & Escamilla, 1997
  • Cymbasoma cheni Lian & Tan, 2019
  • Cymbasoma clairejoanae Suárez-Morales & McKinnon, 2016
  • Cymbasoma claparedei (Giesbrecht, 1892)
  • Cymbasoma clauderazoulsi Suárez-Morales, Goruppi, de Olazabal & Tirelli, 2017
  • Cymbasoma cocoense Suárez-Morales & Morales-Ramírez, 2009
  • Cymbasoma colefaxi Suárez-Morales & McKinnon, 2016
  • Cymbasoma concepcionae Suárez-Morales & Morales Ramírez, 2003
  • Cymbasoma constrictum Suárez-Morales & McKinnon, 2016
  • Cymbasoma dakini Suárez-Morales & McKinnon, 2016
  • Cymbasoma danae (Malaquin, 1896)
  • Cymbasoma davisi Suárez-Morales & Pilz, 2008
  • Cymbasoma fergusoni Suárez-Morales & McKinnon, 2016
  • Cymbasoma filogranarum Malaquin, 1901 (syn. Monstrilla filogranarum (Malaquin, 1901)
  • Cymbasoma frondipes Isaac in Grygier & Suárez-Morales, 2018
  • Cymbasoma galerus Suárez-Morales & McKinnon, 2016
  • Cymbasoma germanicum (Timm, 1893)
  • Cymbasoma ghardaqanum (Isaac, 1975)
  • Cymbasoma gigas (Scott A., 1909)
  • Cymbasoma gracile Gurney, 1927
  • Cymbasoma guererrense Suárez-Morales, 2009
  • Cymbasoma janetae Mageed, 2010
  • Cymbasoma javense Suárez-Morales, in Grygier & Suárez-Morales, 2018 (pro Monstrilla javensis Isaac, 1974a, nomen nudum)
  • Cymbasoma jinigudira Suárez-Morales & McKinnon, 2016
  • Cymbasoma leighrandalli Suárez-Morales & McKinnon, 2016
  • Cymbasoma lenticula Suárez-Morales & McKinnon, 2016
  • Cymbasoma longispinosum (Bourne, 1890)
  • Cymbasoma lourdesae Suárez-Morales & McKinnon, 2016
  • Cymbasoma malaquini (Caullery & Mesnil, 1914)
  • Cymbasoma marioeduardoi Suárez-Morales & McKinnon, 2016
  • Cymbasoma markhasevae Suárez-Morales & McKinnon, 2016
  • Cymbasoma mcalicei Suárez-Morales, 1996
  • Cymbasoma mediterranea Suárez-Morales, Goruppi, de Olazabal & Tirelli, 2017
  • Cymbasoma morii Sekiguchi, 1982
  • Cymbasoma nicolettae Suárez-Morales, 2002
  • Cymbasoma norvegicum Morales & Mercado Salas, 2023 [North Atlantic]
  • Cymbasoma pallidum (Isaac, 1974)
  • Cymbasoma paraconstrictum Suárez-Morales & McKinnon, 2016
  • Cymbasoma pedroi Morales & Mercado Salas, 2023 [North Atlantic]
  • Cymbasoma pseudobidentatum Suárez-Morales, Goruppi, de Olazabal & Tirelli, 2017
  • Cymbasoma pseudoquadridens Suárez-Morales & McKinnon, 2016
  • Cymbasoma quadridens Davis, 1947
  • Cymbasoma quintanarooense (Suárez-Morales, 1994)
  • Cymbasoma rafaelmartinezi Suárez-Morales & McKinnon, 2016
  • Cymbasoma reticulatum (Giesbrecht, 1893)
  • Cymbasoma rigidum Thompson I.C., 1888
  • Cymbasoma rochai Suárez-Morales & Dias, 2001
  • Cymbasoma roscovita (Malaquin, 1901) (syn. Monstrilla roscovita (Malaquin, 1901)
  • Cymbasoma rostratum (Scott T., 1904)
  • Cymbasoma similirostratum (Isaac, 1974)
  • Cymbasoma sinopense Üstün, Terbiyik & Suárez-Morales, 2014
  • Cymbasoma solanderi Suárez-Morales & McKinnon, 2016
  • Cymbasoma specchii Suárez-Morales, Goruppi, de Olazabal & Tirelli, 2017
  • Cymbasoma striatum Isaac, 1974
  • Cymbasoma striifrons Chang, 2012
  • Cymbasoma strzeleckii Suárez-Morales & McKinnon, 2016
  • Cymbasoma tenue (Isaac, 1974) [PDF2]
  • Cymbasoma tergestinum Suárez-Morales, Goruppi, de Olazabal & Tirelli, 2017
  • Cymbasoma tharawalorum Suárez-Morales & McKinnon, 2016
  • Cymbasoma thompsonii (Giesbrecht, 1893)
  • Cymbasoma tirmiziae Khan & Kamaran, 1974
  • Cymbasoma tranteri Suárez-Morales & McKinnon, 2016
  • Cymbasoma tropicum (Wolfenden, 1906)
  • Cymbasoma tumorifrons (Isaac, 1975)
  • Cymbasoma turcorum Suárez-Morales & Ustun, 2018
  • Cymbasoma williamsoni Khan, 1975
  • Cymbasoma zetlandicum (Scott T., 1904)

    THAUMALEUS Kroyer 1849 (?)

  • Thaumaleus bernardensis Wlley, 1920 (?)
  • Thaumaleus frondipes Isaac
  • Thaumaleus germanicus Vervoort, 1988(?)
  • Thaumaleus gigas Scott 1909 (?)

    MAEMONSTRILLA Grygier & Ohtsuka, 2008

  • Maemonstrilla crenulata Suárez-Morales & McKinnon, 2014
  • Maemonstrilla hoi Suárez-Morales & McKinnon, 2014
  • Maemonstrilla hyottoko Grygier & Ohtsuka, 2008
  • Maemonstrilla longipes (Scott A., 1909)
  • Maemonstrilla ohtsukai Suárez-Morales & McKinnon, 2014
  • Maemonstrilla okame Grygier & Ohtsuka, 2008
  • Maemonstrilla polka Grygier & Ohtsuka, 2008
  • Maemonstrilla protuberans Suárez-Morales & McKinnon, 2014
  • Maemonstrilla simplex Grygier & Ohtsuka, 2008
  • Maemonstrilla spinicoxa Grygier & Ohtsuka, 2008
  • Maemonstrilla turgida (Scott A., 1909)

    MONSTRILLA Dana, 1849 (syn. Haemocera Malaquin, 1896; Strilloma Grygier & Suárez-Morlaes, 2018; Thaumaleus Krřyer, 1849;Thaumatoessa Krřyer, 1845; Thaumatoessa Hesse, 1868; Thaumatohessia Giard, 1900

  • Monstrilla anglica Lubbock, 1857
  • Monstrilla angustipes (Isaac, 1974)
  • Monstrilla annulata Suarez Morales, 2024 [Mexico]
  • Monstrilla arctica Davis & Green, 1974 (syn. Caromiobenella arctica (Davis & Green, 1974)
  • Monstrilla armoricana (Hesse, 1868)
  • Monstrilla bahiana Suárez-Morales & Dias, 2001
  • Monstrilla barbata Suárez-Morales & Gasca-Serrano, 1992
  • Monstrilla bernardensis (Willey, 1920) (syn. Monstrillopsis bernardensis (Willey, 1920)
  • Monstrilla boxshalli (Suárez-Morales, 1993) (syn. Cymbasoma boxshalli (Suárez-Morales, 1993)
  • Monstrilla brasiliensis Suárez-Morales & Dias, 2000 (syn. Caromiobenella brasiliensis (Dias & Suárez-Morales, In: Suárez-Morales & Dias, 2000)
  • Monstrilla brevicornis Isaac, 1974
  • Monstrilla bullata (Scott A., 1909) (syn. Cymbasoma bullatum (Scott A., 1909)
  • Monstrilla careli Suárez-Morales & Dias, 2000
  • Monstrilla careloides Suárez-Morales, 2001
  • Monstrilla chetumalensis Suárez-Morales & Castellanos-Osorio, 2014
  • Monstrilla ciqroi (Suárez-Morales, 1993)
  • Monstrilla claparedei (Giesbrecht, 1892) (syn. Cymbasoma claparedei (Giesbrecht, 1892)
  • Monstrilla conjunctiva Giesbrecht, 1902
  • Monstrilla cymbula Scott A., 1909
  • Monstrilla danae Claparčde, 1863 (syn. Thaumaleus thompsoni Giesbrecht, 1893)
  • Monstrilla dubia Scott T., 1904 (syn. Monstrillopsis dubia (Scott T., 1904)
  • Monstrilla elongata Suárez-Morales, 1994
  • Monstrilla filogranarum (Malaquin, 1901)
  • Monstrilla floridana Davis, 1949
  • Monstrilla fosshageni Suárez-Morales & Dias, 2001 (syn. Monstrillopsis fosshageni Suárez-Morales & Dias, 2001)
  • Monstrilla frondipes (Isaac, 1975) (syn. Cymbasoma frondipes Isaac in Grygier & Suárez-Morales, 2018)
  • Monstrilla ghirardellii Suárez-Morales, Goruppi, de Olazabal & Tirelli, 2017
  • Monstrilla germanica (Timm, 1893) (syn. Cymbasoma germanicum (Timm, 1893)
  • Monstrilla ghardaqana (Isaac, 1975) (syn. Cymbasoma ghardaqanum (Isaac, 1975)
  • Monstrilla gibbosa Suárez-Morales & Palomares-García, 1995
  • Monstrilla gigas (Scott A., 1909) (syn. Cymbasoma gigas (Scott A., 1909)
  • Monstrilla globosa Suárez-Morales, 2003
  • Monstrilla gracilicauda Giesbrecht, 1893
  • Monstrilla grandis Giesbrecht, 1891
  • Monstrilla grygieri Suárez-Morales, 2000
  • Monstrilla hamatapex Grygier & Ohtsuka, 1995 (syn. Caromiobenella hamatapex (Grygier & Ohtsuka, 1995)
  • Monstrilla helgolandica Claus, 1863 (syn. Caromiobenella helgolandica (Claus, 1863)
  • Monstrilla humesi Suárez-Morales, 2001
  • Monstrilla ilhoii Lee, J. & Chang, 2016
  • Monstrilla inserta Scott A., 1909
  • Monstrilla investigatoris Sewell, 1949
  • Monstrilla javense Suárez-Morales in Grygier & Suárez-Morales, 2018
  • Monstrilla lata Desai & Bal, 1962
  • Monstrilla leucopis Sars G.O., 1921
  • Monstrilla longa (Isaac, 1974)
  • Monstrilla longicornis Thompson I.C., 1890
  • Monstrilla longipes Scott, 1909 (?)
  • Monstrilla longiremis Giesbrecht, 1893
  • Monstrilla longispinosa Bourne, 1890 (syn. Cymbasoma longispinosum (Bourne, 1890)
  • Monstrilla mahahualensis Suarez Morales 2'22 [Caribbean]
  • Monstrilla malaquini (Caullery & Mesnil, 1914) (syn. Cymbasoma malaquini (Caullery & Mesnil, 1914)
  • Monstrilla mariaeugeniae Suárez-Morales & Islas-Landeros, 1993
  • Monstrilla marioi Suárez-Morales, 2003
  • Monstrilla minuta Isaac, 1975
  • Monstrilla mixta Scott T., 1914
  • Monstrilla nasuta Davis & Green, 1974
  • Monstrilla nicholsii Davis, 1949
  • Monstrilla obesa Isaac, 1975
  • Monstrilla orcula Scott A., 1909
  • Monstrilla pallida (Isaac, 1974) (syn. Cymbasoma pallidum (Isaac, 1974)
  • Monstrilla papilliremis Isaac, 1975
  • Monstrilla pustulata Suárez-Morales & Dias, 2001
  • Monstrilla pygmaea Suárez-Morales, 2000 (syn. Caromiobenella pygmaea (Suárez-Morales, 2000)
  • Monstrilla quintanarooensis (Suárez-Morales, 1994) (syn. Cymbasoma quintanarooense (Suárez-Morales, 1994)
  • Monstrilla quadridens (Davis, 1947) (syn. Cymbasoma quadridens Davis, 1947)
  • Monstrilla rebis Suárez-Morales, 1993
  • Monstrilla reidae Suárez-Morales, 1993
  • Monstrilla reticulata Davis, 1949 (syn. Monstrillopsis reticulata (Davis, 1949)
  • Monstrilla rigida Thompson I.C., 1888 (syn. Cymbasoma rigidum Thompson I.C., 1888)
  • Monstrilla roscovita (Malaquin, 1901)
  • Monstrilla rostrata (Scott T., 1904) (syn. Cymbasoma rostratum (Scott T., 1904)
  • Monstrilla rugosa Davis, 1947
  • Monstrilla satchmoi Suárez-Morales & Dias, 2001
  • Monstrilla scotti Isaac, 1975 in Grygier & Suárez-Morales, 2018
  • Monstrilla similirostrata (Isaac, 1974) (syn. Cymbasoma similirostratum (Isaac, 1974)
  • Monstrilla serricornis Sars G.O., 1921 (syn. Caromiobenella serricornis (Sars G.O., 1921)
  • Monstrilla spinosa Park, 1967 (syn. Spinomonstrilla spinosa (Park, 1967)
  • Monstrilla striata (Isaac, 1974) (syn. Cymbasoma striatum Isaac, 1974)
  • Monstrilla tenuis (Isaac, 1974) (syn. Cymbasoma tenue (Isaac, 1974)
  • Monstrilla tropica (Wolfenden, 1906) (syn. Cymbasoma tropica (Wolfenden, 1906)
  • Monstrilla thompsonii (Giesbrecht, 1892) (syn. Cymbasoma thompsonii (Giesbrecht, 1893)
  • Monstrilla tumorifrons (Isaac, 1974) (syn. Cymbasoma tumorifrons (Isaac, 1975)
  • Monstrilla turgida Scott A., 1909 (syn. Maemonstrilla turgida (Scott A., 1909)
  • Monstrilla typica (Krřyer, 1849)
  • Monstrilla viridis Dana, 1849
  • Monstrilla wandelii Stephensen, 1913
  • Monstrilla xcalakensis Suarez Morales, 2024 [Mexico]
  • Monstrilla zetlandica (Scott T., 1904) (syn. Cymbasoma zetlandicum (Scott T., 1904)

    MONSTRILLOPSIS Sars G.O., 1921

  • Monstrillopsis angustipes Isaac, 1974 (syn. Monstrilla angustipes (Isaac, 1974)
  • Monstrillopsis bernardensis (Willey, 1920)
  • Monstrillopsis boonwurrungorum Surarez-Morales & McKinnon, 2014
  • Monstrillopsis cahuitae Suárez-Morales & Carrillo, 2013
  • Monstrillopsis chathamensis Suárez-Morales & Morales-Ramírez, 2009
  • Monstrillopsis chilensis Suárez-Morales, Bello-Smith & Palma, 2006
  • Monstrillopsis ciqroi Suárez-Morales, 1993 (syn. Monstrilla ciqroi (Suárez-Morales, 1993)
  • Monstrillopsis coreensis Lee, J., D.S. Kim & Chang, 2016
  • Monstrillopsis dubia (Scott T., 1904)
  • Monstrillopsis dubioides Suárez-Morales & Ivanenko, 2004
  • Monstrillopsis ferrarii Suárez-Morales & Ivanenko, 2004
  • Monstrillopsis fosshageni Suárez-Morales & Dias, 2001
  • Monstrillopsis gracile (Gurney, 1927) (syn. Cymbasoma gracile Gurney, 1927
  • Monstrillopsis hastata Surarez-Morales & McKinnon, 2014
  • Monstrillopsis igniterra Suárez-Morales, Ramírez & Derisio, 2008
  • Monstrillopsis longilobata Lee, J., D.S. Kim & Chang, 2016
  • Monstrillopsis nanus Surarez-Morales & McKinnon, 2014
  • Monstrillopsis paradoxa Jeon et al, 2019 [Korea]
  • Monstrillopsis patagonica Morales et al, 2008)
  • Monstrillopsis planifrons Delaforge, Suárez-Morales, Walkusz, Campbell & Mundy, 2017
  • Monstrillopsis pontoeuxinensis Suárez-Morales & Ustun, 2018
  • Monstrillopsis reticulata (Davis, 1949)
  • Monstrillopsis sarsi Isaac, 1975
  • Monstrillopsis zernowi Dolgopol'skaya, 1948

    SPINOMONSTRILLA Suárez-Morales, 2019

  • Spinomonstrilla spinosa (Park, 1967)


    Haemocera Malaquin, 1896 syn. Monstrilla Dana, 1849; Strilloma Grygier & Suárez-Morlaes, 2018 syn.Monstrilla Dana, 1849; Thaumaleus Krřyer, 1849 syn. Monstrilla Dana, 1849 ; Thaumatoessa Krřyer, 1845 syn.Thaumatohessia Giard, 1900; syn. Monstrilla Dana, 1849; Thaumatoessa Hesse, 1868 syn. Thaumatohessia Giard, 1900, accepted as Monstrilla Dana, 1849; Thaumatohessia Giard, 1900 syn. Monstrilla Dana, 1849>


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