syn. Caligoida; Lerneopodidea


CALIGIDAE Burmeister, 1835

This is a large family that includes 31 genera and more than 450 described species (Dojiri & Ho, 2013), most of which are grouped in only two genera (Caligus Mu¨ller, 1785 and Lepeophtheirus Nordmann, 1832) whereas most caligid genera (90%) contain less than ten valid species( Morales Senna et al., 2016)


DARTEVELLIA Brian, 1939

  • Dartevellia bilobata Brian, 1939

    ECHETUS Krřyer, 1863

  • Echetus typicus Krřyer, 1863

    EURYPHORUS Milne Edwards H., 1840 (syn. Arnaeus Krřyer, 1863; Caligeria Dana, 1852; Dysgamus Steenstrup & Lütken, 1861; Elytrophora Gerstaecker, 1853

  • Euryphorus brachypterus (Brachyuptera?) (Gerstaecker, 1853)
  • Euryphorus nordmanni Milne Edwards, 1840
  • Euryphorus suarezi Morales-Serna, Rodriguez-Santiago & Gomez, 2016

    GLOIOPOTES Steenstrup & Lütken, 1861 (syn. Calina Beneden, 1892 )

  • Gloiopotes americanus Cressey, 1967
  • Gloiopotes crassus Wilson & Bere, 1936 (syn. Lepeophtheirus crassus (Wilson & Bere, 1936)
  • Gloiopotes huttoni (Thomson G.M., 1890)
  • Gloiopotes hygomianus Steenstrup & Lütken, 1861
  • Gloiopotes ornatus Wilson C.B., 1905
  • Gloiopotes watsoni Kirtisinghe, 1934

    HERMILIUS Heller, 1865

  • Hermilius alatus Hameed, 1987
  • Hermilius ariodi Prabha & Pillai, 1986
  • Hermilius longicaudus Ho & Kim I.H., 2000
  • Hermilius longicornis Bassett-Smith, 1898
  • Hermilius pseudari Hameed, 1987
  • Hermilius pyriventris Heller, 1865
  • Hermilius tachysuri Pillai & Natarajan, 1977
  • Hermilius youngi Kabata, 1964

    KEY TO SPECIES OF HERMILUS

    KABATAELLA Prabha & Pillai, 1983

  • Kabataella indica Prabha & Pillai, 1984

    LEPEOPHTHEIRUS Nordmann, 1832 (syn. Dentigryps Wilson C.B., 1913; Homoiotes Wilson C.B., 1905 ; Indocaligus Pillai, 1961)

    "Markevich (1940) established Pseudolepeophtheirus Markevich, 1940 for Pseudolepeophtheirus longicauda Markevich, 1940 based on copepods collected from the pleuronectid fish Platichthys stellatus (Pallas, 1787). Dojiri Ho (2013) synonymized the genus and the species with Lepeophtheirus Nordmann, 1832 and Lepeophtheirus parvicruris Fraser, 1920, respectively. Later, Homma et al. (2020) resurrected Markevich's species as a member of Lepeophtheirus, i.e., as L. longicauda (Markevich, 1940)" (from:Kakui & Uyueno, 2020)

  • Lepeophtheirus acutus Heegaard, 1943
  • Lepeophtheirus aesopus Wilson C.B., 1906
  • Lepeophtheirus alvaroi Suárez-Morales & Gasca, 2012
  • Lepeophtheirus anguilli Hameed, 1976
  • Lepeophtheirus anomalus (Pillai, 1967) (syn.s Anuretes anomalus Pillai, 1967
  • Lepeophtheirus appendiculatus Krřyer, 1863
  • Lepeophtheirus argentus Hewitt, 1963
  • Lepeophtheirus atypicus Lin, Ho & Chen, 1996
  • Lepeophtheirus azoricus Özak, Rodrigues, Vieira, Rosa, Yanar, Koyuncu & Boxshall, 2017
  • Lepeophtheirus bagri Dana, 1849
  • Lepeophtheirus bifidus Fraser, 1920
  • Lepeophtheirus bifurcatus Wilson C.B., 1905
  • Lepeophtheirus bonaci Pearse, 1953 [Australia]
  • Lepeophtheirus brachyurus Heller, 1865
  • Lepeophtheirus breviventris Fraser, 1920
  • Lepeophtheirus bychowskyi Gusev, 1951
  • Lepeophtheirus chaenichthyis (Cunningham, 1871)
  • Lepeophtheirus chantoni Gusev, 1951
  • Lepeophtheirus chilensis Wilson C.B., 1905
  • Lepeophtheirus clarionensis Shiino, 1959
  • Lepeophtheirus confusum González, Castro, Muńoz & López, 2016
  • Lepeophtheirus constrictus Wilson C.B., 1908
  • Lepeophtheirus cossyphi Krřyer, 1863
  • Lepeophtheirus crabro Krřyer, 1863
  • Lepeophtheirus crassus (Wilson & Bere, 1936)
  • Lepeophtheirus christianensis Wilson 1944 (?)
  • Lepeophtheirus cuneifer Kabata, 1974
  • Lepeophtheirus curtus (Wilson C.B., 1913)
  • Lepeophtheirus dissimulatus Wilson C.B., 1905
  • Lepeophtheirus distinctus Hewitt, 1963
  • Lepeophtheirus edwardsi Wilson C.B., 1905
  • Lepeophtheirus elegans Gusev, 1951
  • Lepeophtheirus elongatus Heegaard, 1962 (syn. Paralebion elongatus Wilson C.B., 1911
  • Lepeophtheirus eminens Wilson C.B., 1944
  • Lepeophtheirus epinepheli Ho & Dojiri, 1977
  • Lepeophtheirus erecsoni Thomson G.M., 1891
  • Lepeophtheirus etelisi Ho, Chiang & Lin, 2008
  • Lepeophtheirus europaensis Zeddam, Berrebi, Renaud, Raibaut & Gabrion, 1988
  • Lepeophtheirus exilipes Ho & Lin, 2003
  • Lepeophtheirus exsculptus Fischer, 1860
  • Lepeophtheirus fallolunulus Lewis A.G., 1967 (syn. Anuretes fallolunulus (Lewis, A.G., 1967)
  • Lepeophtheirus flores (Beneden, 1892) (syn. Pupulina flores Beneden, 1892
  • Lepeophtheirus formosanus Ho & Lin, 2010
  • Lepeophtheirus frecuens Castro-Romero & Baeza-Kuroki, 1984
  • Lepeophtheirus furcatus (Capart, 1953)
  • Lepeophtheirus goniistii Yamaguti, 1936
  • Lepeophtheirus grohmanni Krřyer, 1863
  • Lepeophtheirus gusevi Moon & Kim I.H., 2012
  • Lepeophtheirus hapalogenyos Yamaguti & Yamasu, 1959
  • Lepeophtheirus hastatus Shiino, 1960
  • Lepeophtheirus heckelii Krřyer, 1863 (syn. Anuretes heckelii (Krřyer, 1863)
  • Lepeophtheirus heegaardi Hewitt, 1963
  • Lepeophtheirus hexagrammi Gusev, 1951
  • Lepeophtheirus hidekoi Ho, 1962
  • Lepeophtheirus hippoglossi (Krřyer, 1837)
  • Lepeophtheirus histiopteridi Kazachenko, Korotaeva & Kurochkin, 1972
  • Lepeophtheirus hospitalis Fraser, 1920
  • Lepeophtheirus hummi Pearse, 1953
  • Lepeophtheirus huttoni Thomson G.M., 1890 (syn. Gloiopotes huttoni (Thomson G.M., 1890)
  • Lepeophtheirus intercurrens Krřyer, 1863
  • Lepeophtheirus interitus Wilson C.B., 1921
  • Lepeophtheirus kabatai Ho & Dojiri, 1977
  • Lepeophtheirus krishnai Kaliyamurthy, 1990
  • Lepeophtheirus lagocephali Pillai, 1963
  • Lepeophtheirus lalandei Kensley & Grindley, 1973
  • Lepeophtheirus lateolabraxi Shen, 1958
  • Lepeophtheirus latigenitalis Pillai & Natarajan, 1977
  • Lepeophtheirus lewisi Hewitt, 1971
  • Lepeophtheirus lichiae Barnard, 1948
  • Lepeophtheirus litus (Lewis A.G., 1964)
  • Lepeophtheirus longiabdominis Shiino, 1960
  • Lepeophtheirus longicauda (Markevich, 1940)
  • Lepeophtheirus longicaudus (Cressey, 1966)
  • Lepeophtheirus longipalpus Bassett-Smith, 1898
  • Lepeophtheirus longipes Wilson C.B., 1905
  • Lepeophtheirus longispinosus Wilson C.B., 1908
  • Lepeophtheirus longiventralis Yü & Wu, 1932
  • Lepeophtheirus marcepes Wilson C.B., 1944
  • Lepeophtheirus marginatus Bere, 1936 (syn.L. christianensis Wilson, 1944; L. orbicularis Shiino, 1960 a)
  • Lepeophtheirus molvae Milne Edwards, 1836
  • Lepeophtheirus monacanthus Heller, 1865
  • Lepeophtheirus mondacola Morales Serna et al.,2023 [Mexico]
  • Lepeophtheirus mugiloidis Villalba & Duran, 1986
  • Lepeophtheirus muraenae Shiino, 1960
  • Lepeophtheirus nanaimoensis Wilson C.B., 1912
  • Lepeophtheirus natalensis Kensley & Grindley, 1973(syn. Caligus furcisetifer Redkar, Rangnekar & Murti, 1949
  • Lepeophtheirus nordmannii (Milne Edwards, 1836)
  • Lepeophtheirus oblitus Kabata, 1973
  • Lepeophtheirus palliatus (Wilson C.B., 1905)
  • Lepeophtheirus paralichthydis Yamaguti & Yamasu, 1960
  • Lepeophtheirus parvicruris Fraser, 1920
  • Lepeophtheirus parviventris Wilson C.B., 1905
  • Lepeophtheirus parvulus Shiino, 1952
  • Lepeophtheirus parvus Wilson C.B., 1908
  • Lepeophtheirus paulus Cressey, 1969
  • Lepeophtheirus pectoralis (Müller O.F., 1776)
  • Lepeophtheirus perpes Leigh-Sharpe, 1934
  • Lepeophtheirus pharaonis (von Nordmann, 1832) (syn. Caligus pharaonis Nordmann, 1832
  • Lepeophtheirus platensis Thomsen, 1949
  • Lepeophtheirus plectropomi Nuńes-Ruivo & Fourmanoir, 1956
  • Lepeophtheirus plotosi Barnard, 1948
  • Lepeophtheirus pollachius Bassett-Smith, 1896
  • Lepeophtheirus polyprioni Hewitt, 1963
  • Lepeophtheirus pravipes Wilson C.B., 1912
  • Lepeophtheirus quadratus Krřyer, 1863
  • Lepeophtheirus remiopsis Dojiri, 1979
  • Lepeophtheirus renalis (Heegaard, 1945)
  • Lepeophtheirus rhinobati Luque, Chaves & Cezar, 1999
  • Lepeophtheirus robertae Boxshall, 2018
  • Lepeophtheirus robustus Krřyer, 1863
  • Lepeophtheirus rotundatus Brian, 1932
  • Lepeophtheirus rotundipes Dojiri, 1979
  • Lepeophtheirus rotundiventris Bassett-Smith, 1898
  • Lepeophtheirus schaadti Passarelli & Tang, 2017
  • Lepeophtheirus scutiger Shiino, 1952
  • Lepeophtheirus sekii Yamaguti, 1936
  • Lepeophtheirus selkirki Atria, 1969
  • Lepeophtheirus semicossyphi Yamaguti, 1939
  • Lepeophtheirus sheni Boxshall & Bellwood, 1981
  • Lepeophtheirus shiinoi Prabha & Pillai, 1986
  • Lepeophtheirus sigani Ho, Kim I.H., Cruz-Lacierda & Nagasawa, 2004
  • Lepeophtheirus simplex Ho, Gómez & Fajer-Avila, 2001
  • Lepeophtheirus spatha Dojiri & Brantley, 1991
  • Lepeophtheirus spinifer Kirtisinghe, 1937
  • Lepeophtheirus sturionis (Krřyer, 1837)
  • Lepeophtheirus suhmi Brady, 1883
  • Lepeophtheirus tamradus Moon & Kim I.H., 2012
  • Lepeophtheirus tenuis (Leidy, 1889)
  • Lepeophtheirus thompsoni Baird, 1850
  • Lepeophtheirus tuberculatus Kim I.H., 1993
  • Lepeophtheirus uluus (ulua?) (Lewis A.G., 1964)
  • Lepeophtheirus unispinosus Pearse, 1953
  • Lepeophtheirus yanezi Stuardo & Fagetti, 1961
  • Lepeophtheirus watanabei Shiino 1954 [Japan](?)
  • Lepeophtheirus zbigniewi Castro-Romero & Baeza-Kuroki, 1981

    MAPPATES Rangnekar, 1958

  • Mappates alter Kabata, 1964
  • Mappates plataxus Rangnekar, 1958

    MARKEVICHUS Özdikmen, 2008 (syn.Caligopsis Markevich, 1940)

  • Markevichus ponticus (Markevich, 1940)

    METACALIGUS Thomsen, 1949

  • Metacaligus latus Ho & Lin, 2002
  • Metacaligus rufus (Wilson C.B., 1908)
  • Metacaligus trichiuri (Krřyer, 1863)
  • Metacaligus unguidentatus (Rangnekar & Murti, 1950)
  • Metacaligus yucatanensis Suárez-Morales, Kim I.H. & Escamilla, 2012

    PARALEBION Wilson C.B., 1911

  • Paralebion aliuncus (Rangnekar, 1955)
  • Paralebion curticaudis Wilson C.B., 1913
  • Paralebion elongatus Wilson C.B., 1911
  • Paralebion pearsei Causey, 1953

    PARAPETALUS Steenstrup & Lütken, 1861

  • Parapetalus caudatus Gnanamuthu 1949 [India] (?)
  • Parapetalus denticulatus (Shen, 1957)(syn. Sinocaligus denticulatus Shen, 1957)
  • Parapetalus dewani Pillai & Hameed, 1987
  • Parapetalus hirsutus (Bassett-Smith, 1898)
  • Parapetalus kurochkini (Kazachenko, 1975) (syn. Caligus kurochkini Kazachenko, 1975
  • Parapetalus longipennatus Rangnekar, 1956
  • Parapetalus occidentalis Wilson C.B., 1908
  • Parapetalus orientalis Steenstrup & Lütken, 1861
  • Parapetalus parapetalopsis (Hameed & Pillai, 1973) (syn. Caligus parapetalopsis Hameed & Pillai, 1973

    PARECHETUS Pillai, 1962

  • Parechetus carangis (Bassett-Smith, 1898)

    PSEUDANURETES Yamaguti, 1936

  • Pseudanuretes anfoozi Adday & Ali, 2020
  • Pseudanuretes chaetodontis Yamaguti, 1936
  • Pseudanuretes chelatus (Prabha & Pillai, 1986)
  • Pseudanuretes fortipedis Kabata, 1965
  • Pseudanuretes indicus Prabha & Pillai, 1986
  • Pseudanuretes papernai Kabata & Deets, 1988
  • Pseudanuretes parvulus (Wilson C.B., 1913)
  • Pseudanuretes pomacanthi Lewis A.G., 1968
  • Pseudanuretes pomacanthodi Prabha & Pillai, 1983
  • Pseudanuretes schmitti Rangnekar, 1957

    PSEUDECHETUS Prabha & Pillai, 1979

  • Pseudechetus fimbriatus Prabha & Pillai, 1979

    PSEUDOPETALUS Pillai, 1962

  • Pseudopetalus caudatus Gnanamuthu, 1950
  • Pseudopetalus dussumieri (Rangnekar, 1957)
  • Pseudopetalus timorensis Izawa, 1995 (syn. Sinocaligus timorensis (Izawa, 1995)

    PUPULINA Beneden, 1892 (syn. Diphyllogaster Brian, 1899)

    "Species of Pupulina Van Beneden, 1892 have been reported from rays of the genera Mobula Rafinesque, 1810 and Manta Bancroft, 1829 from eastern Pacific, the Gulf of Mexico and South Africa (Wilson 1952, Dippenaar and Lebepe 2013). Members of this caligid genus appear to be restricted to species of the ray family Mobulidae (Dojiri and Ho 2013)"

  • Pupulina brevicauda Wilson M.S., 1952
  • Pupulina cliffi Dippenaar & Lebepe, 2013
  • Pupulina flores Beneden, 1892
  • Pupulina keiri Boxshall, 2018
  • Pupulina mantensis Cruz-Quintana, Cańa-Bozada, Suárez-Morales & Santana-Pińeros, 2018
  • Pupulina merira Dippenaar & Lebepe, 2013
  • Pupulina minor Wilson M.S., 1952

    KEY TO SPECIES OF PUPULINA


    SINOCALIGUS Shen, 1957

  • Sinocaligus caudatus (Gnanamuthu, 1950) (syn. Parapetalus caudatus Gnanamuthu, 1950
  • Sinocaligus dussumieri (Rangnekar, 1957) (syn. Pseudopetalus dussumieri (Rangnekar, 1957)
  • Sinocaligus formicoides (Redkar, Rangnekar & Murti, 1949) (syn- Pseudopetalus formicoides Redkar, Rangnekar & Murti, 1949
  • Sinocaligus timorensis (Izawa, 1995)

    SYNESTIUS Steenstrup & Lütken, 1861

  • Synestius caliginus Steenstrup & Lütken, 1861

    TUXOPHORUS Wilson C.B., 1908 (syn. Tuxophoropsis Pillai, 1985 )

  • Tuxophorus caligodes Wilson C.B., 1908
  • Tuxophorus cervicornis Heegaard, 1962
  • Tuxophorus collettei Cressey & Cressey, 1980
  • Tuxophorus cybii Nuńes-Ruivo & Fourmanoir, 1956
  • Tuxophorus tylosuri Rangnekar, 1956 (syn. Caligus tylosuri (Rangnekar, 1956)
  • Tuxophorus wilsoni Kirtisinghe, 1937
  • Tuxophorus zonichthi Uma Devi & Shyamasundari, 1978



    CALVEROCHERIDAE Stock, 1968

    CALVEROCHERES Wilson C.B., 1932 (syn. Calvocheres Wilson C.B., 1932; Echinocheres Hansen, 1902; Hansocheres Wilson C.B., 1924)

  • Calverocheres engeli Stock, 1971
  • Calverocheres globosus (Hansen, 1902)
  • Calverocheres oblongus Stephensen, 1935



    CANCERILLIDAE Giesbrecht, 1897

    CANCERILLA Dalyell, 1851 (syn. Caligidium Claus, 1889)

  • Cancerilla alata Heegaard, 1951
  • Cancerilla ampla Heegaard, 1951
  • Cancerilla durbanensis Stephensen, 1933
  • Cancerilla neozelanica Stephensen, 1927
  • Cancerilla oblonga Bartsch, 1975
  • Cancerilla setifera Bartsch, 1994
  • Cancerilla tubulata Dalyell, 1851
  • Cancerilla vagabunda (Claus, 1889) (syn. Cancerilla tubulata Dalyell, 1851

    CANCERILLOPSIS Stephensen, 1933

  • Cancerillopsis nanaimensis Stephensen, 1933

    OPHIOPSYLLOPSIS Sebastian, 1968

  • Ophiopsyllopsis indicus Sebastian, 1968

    OPHIOPSYLLUS Stock, Humes & Gooding, 1963

  • Ophiopsyllus latus Humes & Hendler, 1999
  • Ophiopsyllus reductus Stock, Humes & Gooding, 1963

    PARARTOTROGUS Scott T. & Scott A., 1893

  • Parartotrogus arcticus Scott T., 1901
  • Parartotrogus minutus Kim 2022 [Korea]
  • Parartotrogus richardi Scott T. & Scott A., 1893

    PARAPHIOPSYLLUS Humes & Hendler, 1972

  • Parophiopsyllus ligatus Humes & Hendler, 1972



    CODOBIDAE Boxshall & Ohtsuka, 2001

    CODOBA Heegaard, 1951

  • Codoba discoveryi Heegaard, 1951



    CORALLIOMYZONTIDAE Humes & Stock, 1991

    CHOLOMYZON Stock & Humes, 1969

  • Cholomyzon brevisetigerum Humes, 1997
  • Cholomyzon multisetum Uyeno & Johnsson, 2018
  • Cholomyzon palpiferum Stock & Humes, 1969
  • Cholomyzon tubastraeae Cheng, Dai & Chang, 2011

    CORALLIOMYZON Humes & Stock, 1991

  • Coralliomyzon latitergum Humes, 1997
  • Coralliomyzon tenens Humes & Stock, 1991

    TEMANUS Humes, 1997

  • Temanus halmaherensis Humes, 1997

    TONDUA Humes, 1997

  • Tondua tholincola Humes, 1997



    DICHELESTHIIDAE Milne Edwards, 1840 (syn. Anthosomatidae Yamaguti, 1963; Anthosomidae Yamaguti, 1963)

    ANTHOSOMA Leach, 1816

  • Anthosoma crassum (Abildgaard, 1794)

    DICHELESTHIUM Hermann, 1804

  • Dichelesthium oblongum (Abildgaard, 1794)



    DICHELINIDAE Boxshall & Ohtsuka, 2001

    DICHELINA Stephensen, 1933

  • Dichelina phormosomae Stephensen, 1933
  • Dichelina seticauda Stock, 1968



    DINOPONTIIDAE Murnane, 1967

    DINOPONTIUS Stock, 1960

  • Dinopontius acuticauda Stock, 1960

    STENOPONTIUS Murnane, 1967

  • Stenopontius boxshalli Malt, 1991
  • Stenopontius humesi Murnane, 1967
  • Stenopontius parvus Boxshall, 1990
  • Stenopontius spinulatus Kim I.H., 2010



    DIRIVULTIDAE Humes & Dojiri, 1981

    Dirivultidae Humes & Dojiri, 1980a is a primitive family of siphonostomatoid copepods whose species have been collected in large numbers from vent fields in the Atlantic and Pacific Oceans (Humes & Segonzac, 1998; Ivanenko & Defaye, 2006a; Gollner et al., 2010; Ivanenko et al., 2011).

    APHOTOPONTIUS Humes, 1987

  • Aphotopontius acanthinus Humes & Lutz, 1994
  • Aphotopontius arcuatus Humes, 1987
  • Aphotopontius atlanteus Humes, 1996
  • Aphotopontius baculigerus Humes, 1987
  • Aphotopontius flexispinus (flexispina in the original description) Humes, 1987
  • Aphotopontius forcipatus Humes, 1987
  • Aphotopontius hydronauticus Humes, 1989
  • Aphotopontius kiost Lee J, Kim D & Kim IH, 2020
  • Aphotopontius limatulus Humes, 1987
  • Aphotopontius mammillatus Humes, 1987
  • Aphotopontius muricatus Lee J, Kim D & Kim IH, 2020
  • Aphotopontius probolus Humes, 1990
  • Aphotopontius rapunculus Humes & Segonzac, 1998 (syn. Rhogobius rapunculus (Humes & Segonzac, 1998)
  • Aphotopontius temperatus Humes 1997 [Azores] (?)

    BENTHOXYNUS Humes, 1984

  • Benthoxynus constrictus Lee J, Kim D & Kim IH, 2020
  • Benthoxynus spiculifer Humes, 1984
  • Benthoxynus tumidiseta Humes, 1989

    CEUTHOECETES Humes & Dojiri, 1980

  • Ceuthoecetes acanthothrix Humes, 1987
  • Ceuthoecetes aliger Humes & Dojiri, 1980
  • Ceuthoecetes cristatus Humes, 1987
  • Ceuthoecetes introversus Humes, 1987

    CHASMATOPONTIUS Humes, 1990

  • Chasmatopontius inflatus Kim S.J. & Kim I.H., 2013
  • Chasmatopontius thescalus Humes, 1990

    DIRIVULTUS Humes & Dojiri, 1981

  • Dirivultus dentaneus Humes & Dojiri, 1981
  • Dirivultus kaiko Uyeno, Kakui, Watanabe & Fujiwara, 2020
  • Dirivultus spinigulatus Humes, 1999

    EXRIMA Humes, 1987

  • Exrima dolichops Humes, 1987
  • Exrima singula Humes, 1987
  • Exrima walteri Ivanenko, Defaye & Ferrari, 2011

    FISSURIOLA Humes, 1987

  • Fissuricola caritus Humes, 1987

    HUMESIPONTIUS Ivanenko & Ferrari, 2003

  • Humesipontius arthuri Ivanenko & Ferrari, 2003

    NILVA Humes, 1987

  • Nilva torifera Humes, 1987

    RHOGOBIUS Humes, 1987

  • Rhogobius contractus Humes, 1987
  • Rhogobius pressulus Humes, 1989
  • Rhogobius rapunculus (Humes & Segonzac, 1998)

    RIMIPONTIUS Humes, 1996

  • Rimipontius mediospinifer Humes, 1996

    SCOTOECETES Humes, 1987

  • Scotoecetes introrsus Humes, 1987

    STYGIOPONTIUS Humes, 1987

  • Stygiopontius appositus Humes, 1989
  • Stygiopontius brevispina Humes, 1991
  • Stygiopontius bulbisetiger Humes, 1996 (syn. Stygiopontius pectinatus Humes, 1987)
  • Stygiopontius cinctiger Humes, 1987
  • Stygiopontius cladarus Humes, 1996
  • Stygiopontius flexus Humes, 1987
  • Stygiopontius geminus Lee J, Kim D & Kim IH, 2020
  • Stygiopontius hispidulus Humes, 1987
  • Stygiopontius horridus Lee J, Kim D & Kim IH, 2020
  • Stygiopontius latulus Humes, 1996
  • Stygiopontius lauensis Humes, 1991
  • Stygiopontius liui Ma, Wang & Li, 2020
  • Stygiopontius lomonosovi Ivanenko & Martínez Arbizu, 2006
  • Stygiopontius lumiger Humes, 1989 (syn. Stygiopontius sentifer Humes, 198)
  • Stygiopontius mirus Humes, 1996
  • Stygiopontius mucroniferus Humes, 1987
  • Stygiopontius paxillifer Humes, 1989
  • Stygiopontius pectinatus Humes, 1987
  • Stygiopontius quadripaxillifer Lee J, Kim D & Kim IH, 2020
  • Stygiopontius quadrispinosus Humes, 1987
  • Stygiopontius regius Humes, 1996
  • Stygiopontius rimivagus Humes, 1997
  • Stygiopontius senckenbergi Ivanenko & Ferrari, 2013
  • Stygiopontius senokuchiae Uyeno, Watanabe & Shimanaga, 2018
  • Stygiopontius sentifer Humes, 1987
  • Stygiopontius serratus Humes, 1996
  • Stygiopontius spinifer Lee J, Kim D & Kim IH, 2020
  • Stygiopontius stabilitus Humes, 1990
  • Stygiopontius teres Humes, 1996
  • Stygiopontius verruculatus Humes, 1987

    KEY TO SPECIES

    KEY FOR THE GENERA OF DIRIVULTIDAE



    DISSONIDAE Kurtz, 1924

    DISSONUS Wilson C.B., 1906 (syn. Caligotrogus Ummerkutty, 1970)

  • Dissonus excavatus Boxshall, Lin, Ho, Ohtsuka, Venmathi Maran & Justine, 2008
  • Dissonus furcatus Kirtisinghe, 1950
  • Dissonus glaber Kurtz, 1924
  • Dissonus heronensis Kabata, 1966
  • Dissonus hoi Tang & Kalman, 2005
  • Dissonus inaequalis Boxshall, Lin, Ho, Ohtsuka, Venmathi Maran & Justine, 2008
  • Dissonus kapuri (Ummerkutty, 1970)
  • Dissonus manteri Kabata, 1966
  • Dissonus nudiventris Kabata, 1965
  • Dissonus ruvetti Nuńes-Ruivo & Fourmanoir, 1956
  • Dissonus pastinum Deets & Dojiri, 1990 (syn. Dissonus nudiventris Kabata, 1965)

  • Dissonus similis Kabata, 1966
  • Dissonus spinifer Wilson C.B., 1906

    Key to species of Dissonus


    INNAPROKOFEVNAS Kazatchenko, 2001

  • Innaprokofevnas orientcolae Kazachenko, 2001



    ECBATHYRIONTIDAE Humes, 1987

    BATHYGORDION Ivanenko & Martinez Arbizu, 2018

  • Bathygordion cliffus Ivanenko & Martinez Arbizu, 2018

    ECBATHYRION Humes, 1987

  • Ecbathyrion prolixicauda Humes, 1987



    ENTOMOLEPIDIDAE Brady, 1899

    Entomolepididae is a small siphonostomatoid family with a distinct shield-like body morphology, comprising 9 genera and 20 species. These genera are divided into two subfamilies, mostly based on their body segmentation, the genera with fused pedigerous somites 2 and 3 are included in the Parmulodinae subfamily, and the genera with free pedigerous somites 2 and 3 are included in the Entomolepinae subfamily (Farias et al., 2024).

    ENTOMOLEPIS Brady, 1899

  • Entomolepis adriae Eiselt, 1959 (syn. Entomopsyllus adriae (Eiselt, 1959)
  • Entomolepis hamondi McKinnon, 1988
  • Entomolepis ovalis Brady, 1899

    ENTOMOPSYLLUS McKinnon, 1988

  • Entomopsyllus adriae (Eiselt, 1959)
  • Entomopsyllus brevicaudatus Lee, J. & I.H. Kim, 2017
  • Entomopsyllus nichollsi McKinnon, 1988
  • Entomopsyllus stocki Kim I.H., 2004
  • Entomopsyllus takara Uyeno & Johnsson, 2018

    LEPEOPSYLLUS Thompson I.C. & Scott A., 1903

  • Lepeopsyllus ovalis Thompson I.C. & Scott A., 1903
  • Lepeopsyllus typicus Thompson I.C. & Scott A., 1903

    PARALEPEOPSYLLUS Ummerkutty, 1960

  • Paralepeopsyllus dambayensis Lee, J. & I.H. Kim, 2017
  • Paralepeopsyllus leei Lee, J. & I.H. Kim, 2017
  • Paralepeopsyllus mannarensis Ummerkutty, 1960

    PARMULELLA Stock, 1992

  • Parmulella emarginata Stock, 1992

    PARMULODES Wilson C.B., 1944

  • Parmulodes verrucosa Wilson C.B., 1944

    Subfamily PARMULODINAE Eiselt, 1959

    NEOPARMULELLA Farias, Neves & Johnsson, 2020

  • Neoparmulella periperiensis Farias, Neves & Johnsson, 2020

    PARMULOPSYLLUS Borges, Farias, Mácola, Neves & Johnsson, 2021

  • Parmulopsyllus iamarinoi Borges, Farias, Mácola, Neves & Johnsson, 2021

    SPONGIOPSYLLUS Johnsson, 2000

  • Spongiopsyllus adventicius Johnsson, 2000
  • Spongiopsyllus aramisi Farias et al. 2024 [Brazil]
  • Spongiopsyllus atypicus Canário, Hurbatah, Rocha, Neves & Johnsson, 2019
  • Spongiopsyllus intermedius Borges, Farias, Mácola, Neves & Johnsson, 2021
  • Spongiopsyllus porthosi Farias et al. 2024 [Brazil]
  • Spongiopsyllus redactus Canario, Neves & Johnsson, 2012

    Key to the species of the Entomolepididae


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