![](sipho1.gif)
syn. Caligoida;
Lerneopodidea CALIGIDAE Burmeister, 1835
Caligidae This is a large
family that includes 31 genera and more than 450
described species (Dojiri & Ho, 2013), most of which
are grouped in only two genera (Caligus Mu¨ller, 1785
and Lepeophtheirus Nordmann, 1832) whereas most
caligid genera (90%) contain less than ten valid
species( Morales Senna et al., 2016)
DARTEVELLIA Brian, 1939
Dartevellia bilobata Brian, 1939
ECHETUS Krřyer, 1863
Echetus typicus Krřyer, 1863
EURYPHORUS Milne Edwards H., 1840 (syn. Arnaeus Krřyer, 1863; Caligeria Dana, 1852; Dysgamus Steenstrup & Lütken, 1861; Elytrophora Gerstaecker, 1853
Euryphorus brachypterus (Brachyuptera?) (Gerstaecker, 1853)
Euryphorus nordmanni Milne Edwards, 1840 ![](pdf.gif)
Euryphorus suarezi Morales-Serna, Rodriguez-Santiago & Gomez, 2016 ![](pdf.gif)
GLOIOPOTES Steenstrup & Lütken, 1861 (syn. Calina Beneden, 1892 )
Gloiopotes americanus Cressey, 1967 ![](pdf.gif)
Gloiopotes crassus Wilson & Bere, 1936 (syn. Lepeophtheirus crassus (Wilson & Bere, 1936)
Gloiopotes huttoni (Thomson G.M., 1890) ![](pdf.gif)
Gloiopotes hygomianus Steenstrup & Lütken, 1861
Gloiopotes ornatus Wilson C.B., 1905
Gloiopotes watsoni Kirtisinghe, 1934
HERMILIUS Heller, 1865
Hermilius alatus Hameed, 1987
Hermilius ariodi Prabha & Pillai, 1986
Hermilius longicaudus Ho & Kim I.H., 2000 ![](pdf.gif)
Hermilius longicornis Bassett-Smith, 1898 ![](pdf.gif)
Hermilius pseudari Hameed, 1987
Hermilius pyriventris Heller, 1865 ![](pdf.gif)
Hermilius tachysuri Pillai & Natarajan, 1977
Hermilius youngi Kabata, 1964
KABATAELLA Prabha & Pillai, 1983
Kabataella indica Prabha & Pillai, 1984
LEPEOPHTHEIRUS Nordmann, 1832 (syn. Dentigryps Wilson C.B. , 1913; Homoiotes Wilson C.B., 1905 ; Indocaligus Pillai, 1961)
"Markevich (1940) established Pseudolepeophtheirus Markevich, 1940 for Pseudolepeophtheirus longicauda Markevich, 1940 based on copepods collected from the pleuronectid fish Platichthys stellatus (Pallas, 1787). Dojiri Ho (2013) synonymized the genus and the species with Lepeophtheirus Nordmann, 1832 and Lepeophtheirus parvicruris Fraser, 1920, respectively. Later, Homma et al. (2020) resurrected Markevich's species as a member of Lepeophtheirus, i.e., as L. longicauda (Markevich, 1940)" (from:Kakui & Uyueno, 2020)
Lepeophtheirus acutus Heegaard, 1943
Lepeophtheirus aesopus Wilson C.B., 1906
Lepeophtheirus alvaroi Suárez-Morales & Gasca, 2012 ![](pdf.gif)
Lepeophtheirus anguilli Hameed, 1976 ![](pdf.gif)
Lepeophtheirus anomalus (Pillai, 1967) (syn.s Anuretes anomalus Pillai, 1967
Lepeophtheirus appendiculatus Krřyer, 1863
Lepeophtheirus argentus Hewitt, 1963
Lepeophtheirus atypicus Lin, Ho & Chen, 1996 ![](pdf.gif)
Lepeophtheirus azoricus Özak, Rodrigues, Vieira, Rosa, Yanar, Koyuncu & Boxshall, 2017 ![](pdf.gif)
Lepeophtheirus bagri Dana, 1849
Lepeophtheirus bifidus Fraser, 1920
Lepeophtheirus bifurcatus Wilson C.B., 1905 ![](pdf.gif)
Lepeophtheirus bonaci Pearse, 1953 [Australia] ![](pdf.gif)
Lepeophtheirus brachyurus Heller, 1865
Lepeophtheirus breviventris Fraser, 1920
Lepeophtheirus bychowskyi Gusev, 1951 ![](pdf.gif)
Lepeophtheirus chaenichthyis (Cunningham, 1871)
Lepeophtheirus chantoni Gusev, 1951
Lepeophtheirus chilensis Wilson C.B., 1905 ![](pdf.gif)
Lepeophtheirus clarionensis Shiino, 1959
Lepeophtheirus confusum González, Castro, Muńoz & López, 2016
Lepeophtheirus constrictus Wilson C.B., 1908
Lepeophtheirus cossyphi Krřyer, 1863
Lepeophtheirus crabro Krřyer, 1863
Lepeophtheirus crassus (Wilson & Bere, 1936)
Lepeophtheirus christianensis Wilson 1944 (?) ![](pdf.gif)
Lepeophtheirus cuneifer Kabata, 1974
Lepeophtheirus curtus (Wilson C.B., 1913)
Lepeophtheirus dissimulatus Wilson C.B., 1905
Lepeophtheirus distinctus Hewitt, 1963
Lepeophtheirus edwardsi Wilson C.B., 1905
Lepeophtheirus elegans Gusev, 1951
Lepeophtheirus elongatus Heegaard, 1962 (syn. Paralebion elongatus Wilson C.B., 1911
Lepeophtheirus eminens Wilson C.B., 1944 ![](pdf.gif)
Lepeophtheirus epinepheli Ho & Dojiri, 1977
Lepeophtheirus erecsoni Thomson G.M., 1891
Lepeophtheirus etelisi Ho, Chiang & Lin, 2008 ![](pdf.gif)
Lepeophtheirus europaensis Zeddam, Berrebi, Renaud, Raibaut & Gabrion, 1988
Lepeophtheirus exilipes Ho & Lin, 2003
![](pdf.gif)
Lepeophtheirus exsculptus Fischer, 1860
Lepeophtheirus fallolunulus Lewis A.G., 1967 (syn. Anuretes fallolunulus (Lewis, A.G., 1967)
Lepeophtheirus flores (Beneden, 1892) (syn. Pupulina flores Beneden, 1892
Lepeophtheirus formosanus Ho & Lin, 2010
Lepeophtheirus frecuens Castro-Romero & Baeza-Kuroki, 1984 ![](pdf.gif)
Lepeophtheirus furcatus (Capart, 1953)
Lepeophtheirus goniistii Yamaguti, 1936
Lepeophtheirus grohmanni Krřyer, 1863
Lepeophtheirus gusevi Moon & Kim I.H., 2012
Lepeophtheirus hapalogenyos Yamaguti & Yamasu, 1959
Lepeophtheirus hastatus Shiino, 1960
Lepeophtheirus heckelii Krřyer, 1863 (syn. Anuretes heckelii (Krřyer, 1863)
Lepeophtheirus heegaardi Hewitt, 1963
Lepeophtheirus hexagrammi Gusev, 1951
Lepeophtheirus hidekoi Ho, 1962
Lepeophtheirus hippoglossi (Krřyer, 1837)
Lepeophtheirus histiopteridi Kazachenko, Korotaeva & Kurochkin, 1972
Lepeophtheirus hospitalis Fraser, 1920
Lepeophtheirus hummi Pearse, 1953
Lepeophtheirus huttoni Thomson G.M., 1890 (syn. Gloiopotes huttoni (Thomson G.M., 1890)
Lepeophtheirus intercurrens Krřyer, 1863
Lepeophtheirus interitus Wilson C.B., 1921
Lepeophtheirus kabatai Ho & Dojiri, 1977
Lepeophtheirus krishnai Kaliyamurthy, 1990
Lepeophtheirus lagocephali Pillai, 1963
Lepeophtheirus lalandei Kensley & Grindley, 1973
Lepeophtheirus lateolabraxi Shen, 1958
Lepeophtheirus latigenitalis Pillai & Natarajan, 1977
Lepeophtheirus lewisi Hewitt, 1971
Lepeophtheirus lichiae Barnard, 1948
Lepeophtheirus litus (Lewis A.G., 1964)
Lepeophtheirus longiabdominis Shiino, 1960
Lepeophtheirus longicauda (Markevich, 1940)
Lepeophtheirus longicaudus (Cressey, 1966)
Lepeophtheirus longipalpus Bassett-Smith, 1898
Lepeophtheirus longipes Wilson C.B., 1905
Lepeophtheirus longispinosus Wilson C.B., 1908
Lepeophtheirus longiventralis Yü & Wu, 1932
Lepeophtheirus marcepes Wilson C.B., 1944 ![](pdf.gif)
Lepeophtheirus marginatus Bere, 1936 (syn.L. christianensis Wilson, 1944; L. orbicularis Shiino, 1960 a)
Lepeophtheirus molvae Milne Edwards, 1836
Lepeophtheirus monacanthus Heller, 1865
Lepeophtheirus mondacola Morales Serna et al.,2023 [Mexico]
Lepeophtheirus mugiloidis Villalba & Duran, 1986 Lepeophtheirus ![](pdf.gif)
Lepeophtheirus muraenae Shiino, 1960
Lepeophtheirus nanaimoensis Wilson C.B., 1912
Lepeophtheirus natalensis Kensley & Grindley, 1973(syn. Caligus furcisetifer Redkar, Rangnekar & Murti, 1949
Lepeophtheirus nordmannii (Milne Edwards, 1836)
Lepeophtheirus oblitus Kabata, 1973
Lepeophtheirus palliatus (Wilson C.B., 1905)
Lepeophtheirus paralichthydis Yamaguti & Yamasu, 1960
Lepeophtheirus parvicruris Fraser, 1920
Lepeophtheirus parviventris Wilson C.B., 1905
Lepeophtheirus parvulus Shiino, 1952
Lepeophtheirus parvus Wilson C.B., 1908
Lepeophtheirus paulus Cressey, 1969 ![](pdf.gif)
Lepeophtheirus pectoralis (Müller O.F., 1776)
Lepeophtheirus perpes Leigh-Sharpe, 1934
Lepeophtheirus pharaonis (von Nordmann, 1832) (syn. Caligus pharaonis Nordmann, 1832
Lepeophtheirus platensis Thomsen, 1949
Lepeophtheirus plectropomi Nuńes-Ruivo & Fourmanoir, 1956
Lepeophtheirus plotosi Barnard, 1948
Lepeophtheirus pollachius Bassett-Smith, 1896
Lepeophtheirus polyprioni Hewitt, 1963
Lepeophtheirus pravipes Wilson C.B., 1912
Lepeophtheirus quadratus Krřyer, 1863
Lepeophtheirus remiopsis Dojiri, 1979
Lepeophtheirus renalis (Heegaard, 1945)
Lepeophtheirus rhinobati Luque, Chaves & Cezar, 1999
Lepeophtheirus robertae Boxshall, 2018
Lepeophtheirus robustus Krřyer, 1863
Lepeophtheirus rotundatus Brian, 1932
Lepeophtheirus rotundipes Dojiri, 1979
Lepeophtheirus rotundiventris Bassett-Smith, 1898
Lepeophtheirus schaadti Passarelli & Tang, 2017 ![](pdf.gif)
Lepeophtheirus scutiger Shiino, 1952
Lepeophtheirus sekii Yamaguti, 1936
Lepeophtheirus selkirki Atria, 1969
Lepeophtheirus semicossyphi Yamaguti, 1939 ![](pdf.gif)
Lepeophtheirus sheni Boxshall & Bellwood, 1981
Lepeophtheirus shiinoi Prabha & Pillai, 1986
Lepeophtheirus sigani Ho, Kim I.H., Cruz-Lacierda & Nagasawa, 2004
Lepeophtheirus simplex Ho, Gómez & Fajer-Avila, 2001 ![](pdf.gif)
Lepeophtheirus spatha Dojiri & Brantley, 1991
Lepeophtheirus spinifer Kirtisinghe, 1937
Lepeophtheirus sturionis (Krřyer, 1837)
Lepeophtheirus suhmi Brady, 1883
Lepeophtheirus tamradus Moon & Kim I.H., 2012
Lepeophtheirus tenuis (Leidy, 1889)
Lepeophtheirus thompsoni Baird, 1850
Lepeophtheirus tuberculatus Kim I.H., 1993
Lepeophtheirus uluus (ulua?) (Lewis A.G., 1964) ![](pdf.gif)
Lepeophtheirus unispinosus Pearse, 1953
Lepeophtheirus yanezi Stuardo & Fagetti, 1961
Lepeophtheirus watanabei Shiino 1954 [Japan](?) ![](pdf.gif)
Lepeophtheirus zbigniewi Castro-Romero & Baeza-Kuroki, 1981
MAPPATES Rangnekar, 1958
Mappates alter Kabata, 1964
Mappates plataxus Rangnekar, 1958
MARKEVICHUS Özdikmen, 2008
Markevichus ponticus (Markevich, 1940)
METACALIGUS Thomsen, 1949
Metacaligus latus Ho & Lin, 2002
Metacaligus rufus (Wilson C.B., 1908)
Metacaligus trichiuri (Krřyer, 1863)
Metacaligus unguidentatus (Rangnekar & Murti, 1950)
Metacaligus yucatanensis Suárez-Morales, Kim I.H. & Escamilla, 2012 ![](pdf.gif)
PARALEBION Wilson C.B., 1911
Paralebion aliuncus (Rangnekar, 1955)
Paralebion curticaudis Wilson C.B., 1913
Paralebion elongatus Wilson C.B., 1911
Paralebion pearsei Causey, 1953
PARAPTALUS Steenstrup & Lütken, 1861
Parapetalus denticulatus (Shen, 1957)(syn. Sinocaligus denticulatus Shen, 1957)
Parapetalus dewani Pillai & Hameed, 1987
Parapetalus hirsutus (Bassett-Smith, 1898)
Parapetalus kurochkini (Kazachenko, 1975) (syn. Caligus kurochkini Kazachenko, 1975
Parapetalus longipennatus Rangnekar, 1956
Parapetalus occidentalis Wilson C.B., 1908
Parapetalus orientalis Steenstrup & Lütken, 1861
Parapetalus parapetalopsis (Hameed & Pillai, 1973) (syn. Caligus parapetalopsis Hameed & Pillai, 1973
PARECHETUS Pillai, 1962
Parechetus carangis (Bassett-Smith, 1898)
PSEUDANURETES Yamaguti, 1936
Pseudanuretes anfoozi Adday & Ali, 2020 ![](pdf.gif)
Pseudanuretes chaetodontis Yamaguti, 1936
Pseudanuretes chelatus (Prabha & Pillai, 1986)
Pseudanuretes fortipedis Kabata, 1965
Pseudanuretes indicus Prabha & Pillai, 1986
Pseudanuretes papernai Kabata & Deets, 1988
Pseudanuretes parvulus (Wilson C.B., 1913)
Pseudanuretes pomacanthi Lewis A.G., 1968
Pseudanuretes pomacanthodi Prabha & Pillai, 1983
Pseudanuretes schmitti Rangnekar, 1957
PSEUDECHETUS Prabha & Pillai, 1979
Pseudechetus fimbriatus Prabha & Pillai, 1979
PSEUDOPETALUS Pillai, 1962
Pseudopetalus caudatus Gnanamuthu, 1950
Pseudopetalus dussumieri (Rangnekar, 1957)
Pseudopetalus timorensis Izawa, 1995 (syn. Sinocaligus timorensis (Izawa, 1995)
PUPULINA Beneden, 1892 (syn. Diphyllogaster Brian, 1899)
"Species of Pupulina Van Beneden, 1892 have been reported from rays of the genera
Mobula Rafinesque, 1810 and Manta Bancroft, 1829 from eastern Pacific, the Gulf of
Mexico and South Africa (Wilson 1952, Dippenaar and Lebepe 2013). Members of
this caligid genus appear to be restricted to species of the ray family Mobulidae (Dojiri
and Ho 2013)"
Pupulina brevicauda Wilson M.S., 1952
Pupulina cliffi Dippenaar & Lebepe, 2013 ![](pdf.gif)
Pupulina flores Beneden, 1892
Pupulina keiri Boxshall, 2018
Pupulina mantensis Cruz-Quintana, Cańa-Bozada, Suárez-Morales & Santana-Pińeros, 2018 ![](pdf.gif)
Pupulina merira Dippenaar & Lebepe, 2013 ![](pdf.gif)
Pupulina minor Wilson M.S., 1952
KEY TO SPECIES OF PUPULINA
SINOCALIGUS Shen, 1957
Sinocaligus caudatus (Gnanamuthu, 1950) (syn. Parapetalus caudatus Gnanamuthu, 1950
Sinocaligus dussumieri (Rangnekar, 1957) (syn. Pseudopetalus dussumieri (Rangnekar, 1957) ![](pdf.gif)
Sinocaligus formicoides (Redkar, Rangnekar & Murti, 1949) (syn- Pseudopetalus formicoides Redkar, Rangnekar & Murti, 1949
Sinocaligus timorensis (Izawa, 1995)
SYNESTIUS Steenstrup & Lütken, 1861
Synestius caliginus Steenstrup & Lütken, 1861
TUXOPHORUS Wilson C.B., 1908 (syn. Tuxophoropsis Pillai, 1985 )
Tuxophorus caligodes Wilson C.B., 1908
Tuxophorus cervicornis Heegaard, 1962
Tuxophorus collettei Cressey & Cressey, 1980
Tuxophorus cybii Nuńes-Ruivo & Fourmanoir, 1956
Tuxophorus tylosuri Rangnekar, 1956 (syn. Caligus tylosuri (Rangnekar, 1956)
Tuxophorus wilsoni Kirtisinghe, 1937
Tuxophorus zonichthi Uma Devi & Shyamasundari, 1978
CALVEROCHERIDAE Stock, 1968
CALVEROCHERES Wilson C.B., 1932 (syn. Calvocheres Wilson C.B., 1932; Echinocheres Hansen, 1902; Hansocheres Wilson C.B., 1924)
Calverocheres engeli Stock, 1971
Calverocheres globosus (Hansen, 1902)
Calverocheres oblongus Stephensen, 1935
CANCERILLIDAE Giesbrecht, 1897
CANCERILLA Dalyell, 1851 (syn. Caligidium Claus, 1889)
Cancerilla alata Heegaard, 1951
Cancerilla ampla Heegaard, 1951
Cancerilla durbanensis Stephensen, 1933
Cancerilla neozelanica Stephensen, 1927
Cancerilla oblonga Bartsch, 1975
Cancerilla setifera Bartsch, 1994
Cancerilla tubulata Dalyell, 1851
Cancerilla vagabunda (Claus, 1889) (syn. Cancerilla tubulata Dalyell, 1851
CANCERILLOPSIS Stephensen, 1933
Cancerillopsis nanaimensis Stephensen, 1933
OPHIOPSYLLOPSIS Sebastian, 1968
Ophiopsyllopsis indicus Sebastian, 1968
OPHIOPSYLLUS Stock, Humes & Gooding, 1963 ![](pdf.gif)
Ophiopsyllus latus Humes & Hendler, 1999
Ophiopsyllus reductus Stock, Humes & Gooding, 1963 ![](pdf.gif)
PARARTOTROGUS Scott T. & Scott A., 1893
Parartotrogus arcticus Scott T., 1901
Parartotrogus minutus Kim 2022 [Korea]
Parartotrogus richardi Scott T. & Scott A., 1893
PARAPHIOPSYLLUS Humes & Hendler, 1972
Parophiopsyllus ligatus Humes & Hendler, 1972
CODOBIDAE Boxshall & Ohtsuka, 2001
CODOBA Heegaard, 1951
Codoba discoveryi Heegaard, 1951
CORALLIOMYZONTIDAE Humes & Stock, 1991 ![](pdf.gif)
CHOLOMYZON Stock & Humes, 1969
Cholomyzon brevisetigerum Humes, 1997
Cholomyzon multisetum Uyeno & Johnsson, 2018 ![](pdf.gif)
Cholomyzon palpiferum Stock & Humes, 1969
Cholomyzon tubastraeae Cheng, Dai & Chang, 2011 ![](pdf.gif)
CORALLIOMYZON Humes & Stock, 1991 ![](pdf.gif)
Coralliomyzon latitergum Humes, 1997
Coralliomyzon tenens Humes & Stock, 1991 ![](pdf.gif)
TEMANUS Humes, 1997
Temanus halmaherensis Humes, 1997
TONDUA Humes, 1997
Tondua tholincola Humes, 1997
DICHELESTHIIDAE Milne Edwards, 1840 (syn. Anthosomatidae Yamaguti, 1963; Anthosomidae Yamaguti, 1963)
ANTHOSOMA Leach, 1816
Anthosoma crassum (Abildgaard, 1794) DICHELESTHIUM Hermann, 1804
Dichelesthium oblongum (Abildgaard, 1794)
DICHELINIDAE Boxshall & Ohtsuka, 2001 ![](pdf.gif)
DICHELINA Stephensen, 1933
Dichelina phormosomae Stephensen, 1933 ![](pdf.gif)
Dichelina seticauda Stock, 1968
DINOPONTIIDAE
Murnane, 1967
DINOPONTIUS Stock, 1960
Dinopontius acuticauda Stock, 1960
STENOPONTIUS Murnane, 1967
Stenopontius boxshalli Malt, 1991 ![](pdf.gif)
Stenopontius humesi Murnane, 1967
Stenopontius parvus Boxshall, 1990
Stenopontius spinulatus Kim I.H., 2010
DIRIVULTIDAE Humes & Dojiri, 1981 ![](pdf.gif)
Dirivultidae Humes & Dojiri, 1980a is a primitive family of
siphonostomatoid copepods whose species have been collected
in large numbers from vent fields in the Atlantic and
Pacific Oceans (Humes & Segonzac, 1998; Ivanenko &
Defaye, 2006a; Gollner et al., 2010; Ivanenko et al., 2011).
APHOTOPONTIUS Humes, 1987 ![](pdf.gif)
Aphotopontius acanthinus Humes & Lutz, 1994
Aphotopontius arcuatus Humes, 1987 ![](pdf.gif)
Aphotopontius atlanteus Humes, 1996
Aphotopontius baculigerus Humes, 1987 ![](pdf.gif)
Aphotopontius flexispinus (flexispina in the original description) Humes, 1987 ![](pdf.gif)
Aphotopontius forcipatus Humes, 1987 ![](pdf.gif)
Aphotopontius hydronauticus Humes, 1989
Aphotopontius kiost Lee J, Kim D & Kim IH, 2020
Aphotopontius limatulus Humes, 1987 ![](pdf.gif)
Aphotopontius mammillatus Humes, 1987 ![](pdf.gif)
Aphotopontius muricatus Lee J, Kim D & Kim IH, 2020 ![](pdf.gif)
Aphotopontius probolus Humes, 1990
Aphotopontius rapunculus Humes & Segonzac, 1998 (syn. Rhogobius rapunculus (Humes & Segonzac, 1998)
BENTHOXYNUS Humes, 1984
Benthoxynus constrictus Lee J, Kim D & Kim IH, 2020 ![](pdf.gif)
Benthoxynus spiculifer Humes, 1984
Benthoxynus tumidiseta Humes, 1989
CEUTHOECETES Humes & Dojiri, 1980
Ceuthoecetes acanthothrix Humes, 1987 ![](pdf.gif)
Ceuthoecetes aliger Humes & Dojiri, 1980
Ceuthoecetes cristatus Humes, 1987 ![](pdf.gif)
Ceuthoecetes introversus Humes, 1987 ![](pdf.gif)
CHASMATOPONTIUS Humes, 1990
Chasmatopontius inflatus Kim S.J. & Kim I.H., 2013
Chasmatopontius thescalus Humes, 1990
DIRIVULTUS Humes & Dojiri, 1981
Dirivultus dentaneus Humes & Dojiri, 1981
Dirivultus kaiko Uyeno, Kakui, Watanabe & Fujiwara, 2020
Dirivultus spinigulatus Humes, 1999
EXRIMA Humes, 1987 ![](pdf.gif)
Exrima dolichops Humes, 1987 ![](pdf.gif)
Exrima singula Humes, 1987 ![](pdf.gif)
Exrima walteri Ivanenko, Defaye & Ferrari, 2011 ![](pdf.gif)
FISSURIOLA Humes, 1987 ![](pdf.gif)
Fissuricola caritus Humes, 1987 ![](pdf.gif)
HUMESIPONTIUS Ivanenko & Ferrari, 2003
Humesipontius arthuri Ivanenko & Ferrari, 2003 ![](pdf.gif)
NILVA Humes, 1987 ![](pdf.gif)
Nilva torifera Humes, 1987 ![](pdf.gif)
RHOGOBIUS Humes, 1987
Rhogobius contractus Humes, 1987 ![](pdf.gif)
Rhogobius pressulus Humes, 1989
Rhogobius rapunculus (Humes & Segonzac, 1998) ![](pdf.gif)
RIMIPONTIUS Humes, 1996
Rimipontius mediospinifer Humes, 1996
SCOTOECETES Humes, 1987 ![](pdf.gif)
Scotoecetes introrsus Humes, 1987 ![](pdf.gif)
STYGIOPONTIUS Humes, 1987 ![](pdf.gif)
Stygiopontius appositus Humes, 1989 ![](pdf.gif)
Stygiopontius brevispina Humes, 1991 ![](pdf.gif)
Stygiopontius bulbisetiger Humes, 1996 (syn. Stygiopontius pectinatus Humes, 1987)
Stygiopontius cinctiger Humes, 1987 ![](pdf.gif)
Stygiopontius cladarus Humes, 1996
Stygiopontius flexus Humes, 1987 ![](pdf.gif)
Stygiopontius geminus Lee J, Kim D & Kim IH, 2020 ![](pdf.gif)
Stygiopontius hispidulus Humes, 1987 ![](pdf.gif)
Stygiopontius horridus Lee J, Kim D & Kim IH, 2020 ![](pdf.gif)
Stygiopontius latulus Humes, 1996
Stygiopontius lauensis Humes, 1991 ![](pdf.gif)
Stygiopontius liui Ma, Wang & Li, 2020
Stygiopontius lomonosovi Ivanenko & Martínez Arbizu, 2006 ![](pdf.gif)
Stygiopontius lumiger Humes, 1989 (syn. Stygiopontius sentifer Humes, 198) 7
Stygiopontius mirus Humes, 1996
Stygiopontius mucroniferus Humes, 1987 ![](pdf.gif)
Stygiopontius paxillifer Humes, 1989 ![](pdf.gif)
Stygiopontius pectinatus Humes, 1987 ![](pdf.gif)
Stygiopontius quadripaxillifer Lee J, Kim D & Kim IH, 2020 ![](pdf.gif)
Stygiopontius quadrispinosus Humes, 1987 ![](pdf.gif)
Stygiopontius regius Humes, 1996
Stygiopontius rimivagus Humes, 1997
Stygiopontius senckenbergi Ivanenko & Ferrari, 2013
Stygiopontius senokuchiae Uyeno, Watanabe & Shimanaga, 2018 ![](pdf.gif)
Stygiopontius sentifer Humes, 1987 ![](pdf.gif)
Stygiopontius serratus Humes, 1996
Stygiopontius spinifer Lee J, Kim D & Kim IH, 2020 ![](pdf.gif)
Stygiopontius stabilitus Humes, 1990
Stygiopontius teres Humes, 1996
Stygiopontius verruculatus Humes, 1987 ![](pdf.gif)
KEY TO SPECIES
KEY FOR THE GENERA OF DIRIVULTIDAE
DISSONIDAE Kurtz, 1924
DISSONUS Wilson C.B., 1906 (syn. Caligotrogus Ummerkutty, 1970)
Dissonus excavatus Boxshall, Lin, Ho, Ohtsuka, Venmathi Maran & Justine, 2008
Dissonus furcatus Kirtisinghe, 1950
Dissonus glaber Kurtz, 1924
Dissonus heronensis Kabata, 1966 ![](pdf.gif)
Dissonus hoi Tang & Kalman, 2005
Dissonus inaequalis Boxshall, Lin, Ho, Ohtsuka, Venmathi Maran & Justine, 2008
Dissonus kapuri (Ummerkutty, 1970)
Dissonus manteri Kabata, 1966
Dissonus nudiventris Kabata, 1965
Dissonus ruvetti Nuńes-Ruivo & Fourmanoir, 1956
Dissonus pastinum Deets & Dojiri, 1990 (syn. Dissonus nudiventris Kabata, 1965)
Dissonus similis Kabata, 1966 ![](pdf.gif)
Dissonus spinifer Wilson C.B., 1906
Key to species of Dissonus ![](pdf.gif)
INNAPROKOFEVNAS Kazatchenko, 2001
Innaprokofevnas orientcolae Kazachenko, 2001
ECBATHYRIONTIDAE Humes, 1987 ![](pdf.gif)
BATHYGORDION Ivanenko & Martinez Arbizu, 2018 ![](pdf.gif)
Bathygordion cliffus Ivanenko & Martinez Arbizu, 2018 ![](pdf.gif)
ECBATHYRION Humes, 1987
Ecbathyrion prolixicauda Humes, 1987 ![](pdf.gif)
ENTOMOLEPIDIDAE Brady, 1899
ENTOMOLEPIS Brady, 1899
Entomolepis adriae Eiselt, 1959 (syn. Entomopsyllus adriae (Eiselt, 1959)
Entomolepis hamondi McKinnon, 1988
Entomolepis ovalis Brady, 1899
ENTOMOPSYLLUS McKinnon, 1988
Entomopsyllus adriae (Eiselt, 1959)
Entomopsyllus brevicaudatus Lee, J. & I.H. Kim, 2017 ![](pdf.gif)
Entomopsyllus nichollsi McKinnon, 1988
Entomopsyllus stocki Kim I.H., 2004 ![](pdf.gif)
Entomopsyllus takara Uyeno & Johnsson, 2018 ![](pdf.gif)
LEPEOPSYLLUS Thompson I.C. & Scott A., 1903
Lepeopsyllus ovalis Thompson I.C. & Scott A., 1903
Lepeopsyllus typicus Thompson I.C. & Scott A., 1903
PARALEPEOPSYLLUS Ummerkutty, 1960
Paralepeopsyllus dambayensis Lee, J. & I.H. Kim, 2017
Paralepeopsyllus leei Lee, J. & I.H. Kim, 2017
Paralepeopsyllus mannarensis Ummerkutty, 1960
PARMULELLA Stock, 1992 ![](pdf.gif)
Parmulella emarginata Stock, 1992
PARMULODES Wilson C.B., 1944
Parmulodes verrucosa Wilson C.B., 1944 Subfamily PARMULODINAE Eiselt, 1959
NEOPARMULELLA Farias, Neves & Johnsson, 2020 ![](pdf.gif)
Neoparmulella periperiensis Farias, Neves & Johnsson, 2020 ![](pdf.gif)
PARMULOPSYLLUS Borges, Farias, Mácola, Neves & Johnsson, 2021 ![](pdf.gif)
Parmulopsyllus iamarinoi Borges, Farias, Mácola, Neves & Johnsson, 2021 ![](pdf.gif)
SPONGIOPSYLLUS Johnsson, 2000 ![](pdf.gif)
Spongiopsyllus adventicius Johnsson, 2000 ![](pdf.gif)
Spongiopsyllus atypicus Canário, Hurbatah, Rocha, Neves & Johnsson, 2019 ![](pdf.gif)
Spongiopsyllus intermedius Borges, Farias, Mácola, Neves & Johnsson, 2021 ![](pdf.gif)
Spongiopsyllus redactus Canario, Neves & Johnsson, 2012 ![](pdf.gif)
Key to the species of the Entomolepididae
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