EUDACTYLINIDAE Wilson C.B., 1932

"The copepod parasite family Eudactylinidae Wilson, 1932 comprises 12 genera with 46 valid species and is characterised by elongated cylindrical body demarcated into cephalothorax and an unsegmented trunk except in the genera Eudactylina Van Beneden, 1853, Nemesis Risso, 1826 and Kroyeria Van Beneden, 1853 (Pillai 1985; Walter and Boxshall 2018). Seven of these genera are monotypic and most of them are host specific (Walter and Boxshall 2018), infesting the gills of elasmobranchs and rarely on teleost like Lichia amia Linnaeus"

BARIAKA Cressey, 1966

  • Bariaka alopiae Cressey, 1966
  • Bariaka pamelae (Laubier, Maillard & Oliver, 1966) (syn. Protodactylina pamelae Laubier, Maillard & Oliver, 1966

    CARNIFOSSORIUS Deets & Ho, 1988

  • Carnifossorius siamensis Deets & Ho, 1988

    DANGOKA Izawa, 2011

  • Dangoka japonica Izawa, 2011

    EUDACTYLINA van Beneden, 1853

  • Eudactylina acanthii Scott A., 1901
  • Eudactylina acuta van Beneden, 1853
  • Eudactylina alata Pillai, 1968
  • Eudactylina aspera Heller, 1865
  • Eudactylina bicornis Hameed, Kumar & Mathew, 1990
  • Eudactylina breviabdomina Pearse, 1952
  • Eudactylina brevicauda Hameed, Kumar & Mathew, 1990
  • Eudactylina chelata Hameed, Kumar & Mathew, 1990
  • Eudactylina chilensis Ho & McKinney, 1981
  • Eudactylina corrugata Bere, 1930
  • Eudactylina dasyati Izawa, 2011
  • Eudactylina dollfusi Brian, 1924
  • Eudactylina gymnuri Izawa, 2011
  • Eudactylina indivisa Castro-Romero & Baeza-Kuroki, 1991
  • Eudactylina lancifera Pillai, 1968
  • Eudactylina leptochariae Diebakate & Raibaut, 2000
  • Eudactylina longispina Bere, 1936
  • Eudactylina minuta Scott T., 1904
  • Eudactylina mobuli Hameed, Kumar & Mathew, 1990
  • Eudactylina musteli Izawa, 2011
  • Eudactylina myliobatidos Luque & Farfán, 1991
  • Eudactylina niger (Wilson CB, 1905) (syn. Eudactylinodes niger (Wilson C.B., 1905)
  • Eudactylina nigra Wilson C.B., 1905 (syn. Eudactylinodes niger (Wilson C.B., 1905)
  • Eudactylina oliveri Laubier, 1968
  • Eudactylina papillosa Kabata, 1970
  • Eudactylina parva Castro-Romero & Baeza-Kuroki, 1991
  • Eudactylina peruensis Luque & Farfan, 1991
  • Eudactylina pollex Cressey, 1967
  • Eudactylina puriensis Tripathi, 1962
  • Eudactylina pusilla Cressey, 1967
  • Eudactylina rachelae Green 1958 [British waters]
  • Eudactylina rhinobati Raibaut & Essafi, 1979
  • Eudactylina similis Scott T., 1902
  • Eudactylina spinula Pearse, 1950
  • Eudactylina squamosa Bere, 1936
  • Eudactylina squamosa Bere, 1936
  • Eudactylina taeniuri Izawa, 2011
  • Eudactylina tuberifera Castro-Romero & Baeza-Kuroki, 1987
  • Eudactylina turgipes Bere, 1936
  • Eudactylina uncinata Wilson C.B., 1908 (syn. Eudactylinodes niger (Wilson C.B., 1905)
  • Eudactylina valei Nuñes-Ruivo, 1956
  • Eudactylina vilelai Nuñes-Ruivo, 1956

    EUDACTYLINELLA Wilson C.B., 1932

  • Eudactylinella alba Wilson C.B., 1932

    EUDACTYLINODES Willson C.B., 1932

  • Eudactylinodes keratophagus Deets & Benz, 1986
  • Eudactylinodes niger (Wilson C.B., 1905)
  • Eudactylinodes uncinata (Wilson C.B., 1908) (syn. Eudactylinodes niger (Wilson C.B., 1905)

    EUDACTYLINOPSIS Pillai, 1968

  • Eudactylinopsis curvatus Pillai, 1968

    HETEROCLADIUS Deets & Ho, 1988

  • Heterocladius abyssetes Deets & Ho, 1988

    JANINECAIRA Benz, Smith, Bullard & Braswell, 2007

  • Janinecaira darkthread Benz, Smith, Bullard & Braswell, 2007

    JUSHEYUS Deets & Benz, 1987

  • Jusheyus shogunus Deets & Benz, 1987

    NEMESIS Risso, 1826 (syn. Ergasilina Beneden, 1851; Pagodina Beneden, 1853 )

  • Nemesis aggregatus Cressey, 1967
  • Nemesis atlantica Wilson C.B., 1922
  • Nemesis carchariaeglauci (Hesse, 1883)
  • Nemesis lamna Risso, 1826 (syn. Nemesis lamna lamna Risso, 1826)
  • Nemesis macrocephalus Shiino, 1957
  • Nemesis pallida Wilson C.B., 1932
  • Nemesis pilosus Pearse, 1951[Bahamas]
  • Nemesis robusta (Beneden, 1851)
  • Nemesis santhadevii Maran et al., 2022 [ Malaysia]
  • Nemesis sphyrnae Rangnekar, 1984
  • Nemesis spinulosus Cressey, 1970
  • Nemesis tiburo Pearse, 1952
  • Nemesis vermi Scott A., 1929
  • Nemesis versicolor Wilson C.B., 1913

    PROTODACTYLINA Laubier, Maillard & Oliver, 1966

  • Protodactylina pamelae Laubier, Maillard & Oliver, 1966



    HATSCHEKIIDAE Kabata, 1979

    BASSETTITHIA Wilson C.B., 1922

  • Bassettithia congri (Stebbing, 1900)

    BRACHIHATSCHEKIA Castro-Romero & Baeza-Kuroki, 1989

  • Brachihatschekia lobulata Castro-Romero & Baeza-Kuroki, 1989

    CONGERICOLA Beneden, 1854 (syn. Cycnus Milne Edwards H., 1840; Metahatschekia Yamaguti & Yamasu, 1959)

  • Congericola congeri (Yamaguti & Yamasu, 1959)
  • Congericola gracilis (Milne Edwards, 1840)
  • Congericola kabatai Hewitt, 1975
  • Congericola labrimixti (Hesse, 1878) (syn. Hatschekia labracis (Beneden, 1871)
  • Congericola pallida Beneden, 1854
  • Congericola pagellibogueravei (Hesse, 1878) accepted as Hatschekia pagellibogneravi (Hesse, 1878)

    HATSCHEKIA Poche, 1902 (syn. Caetrodes Wilson C.B., 1906; Pseudoclavella Bassett-Smith, 1898)

    "Hatschekia Poche, 1902, is one of the major genera of Copepoda parasitic on actinopterygian fishes (Boxshall & Halsey 2004). In 1985, Jones recognized 68 species as valid. Since then 41 new species have been described as new (Castro & Baeza 1986; El-Rashidy & Boxshall 2011; Ho & Kim 2001; Jones & Cabral 1990; Kabata 1991; Pillai 1985; Uyeno & Nagasawa 2009b, 2010a–c; Villalba 1986;). Currently, in addition to the shape of the dorsal frame of the cephalothorax, the ratios of body parts and appendages have been established as valuable characters to distinguish morphologically similar species in the genus (Uyeno & Nagasawa 2009b, 2010a–c). Several peculiar morphological features (i.e., parabasal papilla and rostral process) also have been used to identify Hatschekia species.

  • Hatschekia affluens Castro-Romero & Baeza-Kuroki, 1986
  • Hatschekia aigoi Izawa, 2016
  • Hatschekia albirubra Wilson C.B., 1913
  • Hatschekia amphiprocessa Castro-Romero & Baeza-Kuroki, 1986
  • Hatschekia amplicapa Pearse, 1951[Bahamas]
  • Hatschekia angulata Pearse, 1951 [Bahamas]
  • Hatschekia anomalis Izawa, 2015
  • Hatschekia argyops Uma Devi & Shyamasundari, 1980
  • Hatschekia atagonel Jones J.B., 1985
  • Hatschekia aulacocephalis Izawa, 2016
  • Hatschekia awatati Yamaguti, 1939 (syn. Prohatschekia awatati (Yamaguti, 1939)
  • Hatschekia balistae Nuñes-Ruivo, 1954
  • Hatschekia becuni Jones J.B. & Cabral, 1990
  • Hatschekia bibullae Uyeno & Nagasawa, 2010
  • Hatschekia bicaudata Kabata, 1991
  • Hatschekia bifurcata Yamaguti & Yamasu, 1959
  • Hatschekia bodiani Nuñes-Ruivo, 1954
  • Hatschekia boonah Uyeno & Nagasawa, 2013
  • Hatschekia bougisi Nuñes-Ruivo, 1954
  • Hatschekia bramae van Beneden, 1870
  • Hatschekia branchiostegi Yamaguti, 1939
  • Hatschekia breviramus Lewis A.G., 1967
  • Hatschekia brotulae Nuñes-Ruivo, 1954
  • Hatschekia cadenati Nuñes-Ruivo, 1954
  • Hatschekia caudata Pillai, 1985
  • Hatschekia cepolae Yamaguti, 1939
  • Hatschekia cernae Goggio, 1905
  • Hatschekia churaumi Uyeno & Nagasawa, 2010
  • Hatschekia cirrhitichthysicola Izawa, 2016
  • Hatschekia clava Kabata, 1991
  • Hatschekia clavata (Müller O.F., 1776)
  • Hatschekia cluthae (Scott T., 1902)
  • Hatschekia conifer Cressey, 1970
  • Hatschekia conifera Yamaguti, 1939
  • Hatschekia couardi Nuñes-Ruivo, 1954
  • Hatschekia crenata Hewitt, 1969
  • Hatschekia crenulata Kabata, 1991
  • Hatschekia curvata Yamaguti & Yamasu, 1959
  • Hatschekia cyanopodus Lee, S., Lee, W. & Boxshall, 2013
  • Hatschekia cylindrica Shiino, 1957
  • Hatschekia cylindrus Uyeno & Nagasawa, 2009
  • Hatschekia delamarei Nuñes-Ruivo, 1954
  • Hatschekia difficilis Kabata, 1991
  • Hatschekia elegans Kabata, 1991
  • Hatschekia ellipsocorpa Uyeno & Nagasawa, 2013
  • Hatschekia elliptica Pillai, 1968
  • Hatschekia elongata Redkar, Rangnekar & Murti, 1950
  • Hatschekia etelisicola Lee, S., Lee, W. & Boxshall, 2013
  • Hatschekia euanus Lee, S., Lee, W. & Boxshall, 2013
  • Hatschekia exigua Pearse, 1951[Bahamas]
  • Hatschekia flatti Uma Devi & Shyamasundari, 1980
  • Hatschekia foliolata Redkar, Rangnekar & Murti, 1950
  • Hatschekia fukurubi Uyeno & Nagasawa, 2010
  • Hatschekia fuscoguttatus Lee, S., Lee, W. & Boxshall, 2013
  • Hatschekia fusiformis Shiino, 1957
  • Hatschekia geniculata Uyeno & Nagasawa, 2013
  • Hatschekia gerro Leigh-Sharpe, 1936
  • Hatschekia girelli Jones J.B. & Cabral, 1990
  • Hatschekia gracilis Yamaguti, 1954
  • Hatschekia gymnocranii Izawa, 2016
  • Hatschekia hanguyenvani Kazachenko, Kovaleva, Nguyen & Ngo, 2017
  • Hatschekia harkema Pearse, 1948
  • Hatschekia hemicyclium Uyeno & Nagasawa, 2010
  • Hatschekia hemigymni Kabata, 1991
  • Hatschekia hippoglossi (Cuvier, 1830)
  • Hatschekia hoplobrotulae Izawa, 2015
  • Hatschekia iniisticola Izawa, 2016
  • Hatschekia insolita Wilson C.B., 1913
  • Hatschekia iridescens Wilson C.B., 1913
  • Hatschekia ischnon Leigh-Sharpe, 1936
  • Hatschekia izenaensis Uyeno & Nagasawa, 2010
  • Hatschekia japonica Jones J.B., 1985
  • Hatschekia jejuensis Moon & Kim I.H., 2013
  • Hatschekia jonesi Uyeno & Nagasawa, 2010
  • Hatschekia jorgei Nuñes-Ruivo, 1954
  • Hatschekia kabatai Uyeno & Nagasawa, 2010
  • Hatschekia khahajya Uyeno & Nagasawa, 2010
  • Hatschekia kuroshioensis Uyeno & Nagasawa, 2010
  • Hatschekia labracis (Beneden, 1871)
  • Hatschekia laeopsi Izawa, 2018
  • Hatschekia legouili Nuñes-Ruivo, 1954
  • Hatschekia leptoscari Yamaguti, 1939
  • Hatschekia lima Uyeno & Nagasawa, 2009
  • Hatschekia linearis Wilson C.B., 1913
  • Hatschekia longiabdominalis Uyeno & Nagasawa, 2013
  • Hatschekia longibrachium Yamaguti, 1939
  • Hatschekia longigenitalis Yamaguti, 1954
  • Hatschekia louti Lee, S., Lee, W. & Boxshall, 2013
  • Hatschekia maculatus Lee, S., Lee, W. & Boxshall, 2013
  • Hatschekia manea Jones J.B. & Cabral, 1990
  • Hatschekia megacephala Kabata, 1991
  • Hatschekia mihkagan Uyeno & Nagasawa, 2010
  • Hatschekia modesta Kabata, 1965
  • Hatschekia monacanthi Yamaguti, 1939
  • Hatschekia mongarah Uyeno & Nagasawa, 2010
  • Hatschekia mulli (van Beneden, 1851)
  • Hatschekia multibarbatae Izawa, 2015
  • Hatschekia nagasawai Paschoal, 2022
  • Hatschekia nahaensis Yamaguti, 1953
  • Hatschekia nakamurai Uyeno & Nagasawa, 2010
  • Hatschekia napoleoni Jones J.B. & Cabral, 1990
  • Hatschekia nemipteri Izawa, 2016
  • Hatschekia neobythitesi Izawa, 2015
  • Hatschekia niger Lee, S., Lee, W. & Boxshall, 2013
  • Hatschekia nodosa Ho & Kim I.H., 2001
  • Hatschekia nohu Villalba, 1986
  • Hatschekia novaculopsi Izawa, 2016
  • Hatschekia oblonga Wilson C.B., 1913
  • Hatschekia ostracii Yamaguti, 1953
  • Hatschekia ovalis (Bassett-Smith, 1898)
  • Hatschekia pacifica Cressey, 1970
  • Hatschekia pagellibogueravei (Hesse, 1878)
  • Hatschekia pagrosomi Yamaguti, 1939
  • Hatschekia papillifera Kabata, 1991
  • Hatschekia parva Pearse, 1951[Bahamas]
  • Hatschekia paracaesioi Izawa, 2016
  • Hatschekia petiti Nuñes-Ruivo, 1954
  • Hatschekia pholas (Wilson C.B., 1906) (syn. Caetrodes pholas Wilson, 1906)
  • Hatschekia pinguis Wilson C.B., 1908
  • Hatschekia plectopomi Ho & Dojiri, 1978
  • Hatschekia pontini Nuñes-Ruivo, 1954
  • Hatschekia priacanthis Izawa, 2016
  • Hatschekia prionoti Pearse, 1947
  • Hatschekia pseudobalistesi Uyeno & Nagasawa, 2010
  • Hatschekia pseudolabri Yamaguti, 1953
  • Hatschekia pseudostracii Uyeno & Nagasawa, 2010
  • Hatschekia pygmaea Scott T. & Scott A., 1913
  • Hatschekia quadrabdominalis Yü, 1933
  • Hatschekia quadrata Hewitt, 1969
  • Hatschekia rhombocephalis Izawa, 2016
  • Hatschekia rohdei Kabata, 1991
  • Hatschekia rotundigenitalis Yamaguti, 1939
  • Hatschekia sargi Yamaguti, 1939
  • Hatschekia scari (Krøyer, 1838)
  • Hatschekia sebastisci Yamaguti, 1939 (syn. Prohatschekia sebastisci (Yamaguti, 1939)
  • Hatschekia serrana Pearse, 1952 [Australia] (?)
  • Hatschekia seyi Ho & Kim I.H., 2001
  • Hatschekia shari Uyeno & Ali, 2013
  • Hatschekia sigani Uma Devi & Shyamasundari, 1980
  • Hatschekia siganicola El-Rashidy & Boxshall, 2011
  • Hatschekia sphyraeni Pillai, 1960
  • Hatschekia squamata Jones J.B. & Cabral, 1990
  • Hatschekia squamigera Kabata, 1991
  • Hatschekia subpinguis Brian, 1913
  • Hatschekia sumireyakko Uyeno & Nagasawa, 2012
  • Hatschekia sunaoi Uyeno & Nagasawa, 2009
  • Hatschekia synagris Yamaguti, 1954
  • Hatschekia tanysoma Ho & Kim I.H., 2001
  • Hatschekia tenuis (Heller, 1865)
  • Hatschekia teuthidis Yamaguti, 1954
  • Hatschekia thompsoni Heegaard, 1964
  • Hatschekia thyrsitoidesi Izawa, 2016
  • Hatschekia triannuli Uyeno & Nagasawa, 2012
  • Hatschekia uncata Wilson C.B., 1913
  • Hatschekia zanpa Uyeno & Nagasawa, 2010

    Key to females of the valid species of Hatschekia parasitic on pufferfishes


    LAMINOHATSCHEKIA Boxshall, 1989

  • Laminohatschekia synaphobranchi Boxshall, 1989

    MIHBAICOLA Uyeno, 2013

  • Mihbaicola sakamakii Uyeno, 2013

    PROHATSCHEKIA Nunes-Ruivo, 1954

    "Prohatschekia was established by Nunes-Ruivo (1954) to accommodate hatschekiids with three pairs of biramous swimming legs. It was treated as a valid genus by Kabata (1979) and by Boxshall and Halsey (2004). There are currently 8 described species" (from: Hamza et al., 2017)

  • Prohatschekia antennalis Avdeev & Kazachenko, 1985
  • Prohatschekia awatati (Yamaguti, 1939)
  • Prohatschekia cremouxi Nunes-Ruivo, 1954
  • Prohatschekia laguncula Shiino, 1957
  • Prohatschekia mediterranea Hamza, Boxshall & Kechemir-Issad, 2007
  • Prohatschekia pseudocremouxi Moon, Youn & Venmathi Maran, 2016
  • Prohatschekia sebastisci (Yamaguti, 1939)
  • Prohatschekia stocki Kabata, 1990

    KEY TO SPECIES OF PROHATSCHEKIA


    PSEUDOCONGERICOLA Yü, 1933

  • Pseudocongericola chefoonensis Yü, 1933

    WYNNOWENIA Boxshall, 1987

  • Wynnowenia distinctus Boxshall, 1987


    Caetrodes Wilson C.B., 1906 syn. Hatschekia Poche, 1902; Cycnus Milne Edwards H., 1840 accepted as Congericola Beneden, 1854; Metahatschekia Yamaguti & Yamasu, 1959 syn.s Congericola Beneden, 1854; Pseudoclavella Bassett-Smith, 1898 (syn. Hatschekia Poche, 1902



    HYPONEOIDAE Heegaard, 1962

    GREENIEDEETS Benz, 2006

  • Greeniedeets buccaneer Benz, 2006

    HYPONEO Heegaard, 1962

  • Hyponeo australis Heegaard, 1962

    TAUTOCHONDRIA Ho, 1987

  • Tautochondria dolichoura Ho, 1987
  • Tautochondria magellanica Muñoz, Hernández & López, 2023



    KROYERIIDAE Kabata, 1979

    KROEYERINA Wilson C.B., 1932

  • Kroeyerina benzorum Deets, 1987
  • Kroeyerina cortezensis Deets, 1987
  • Kroeyerina deborahae Deets, 1987
  • Kroeyerina deetsorum Benz, Smith & Bullard, 2001
  • Kroeyerina elongata Wilson C.B., 1932
  • Kroeyerina mobulae Deets, 1987
  • Kroeyerina nasuta Wilson C.B., 1932
  • Kroeyerina scottorum Cressey, 1970
  • Kroeyerina sudamericana Irigoitia, Cantatore, Incorvaia & Timi, 2016
  • Kroeyerina meridionalis Ramirez, 1975 accepted as Prokroyeria meridionalis (Ramirez, 1975)

    KROYERIA Beneden, 1853 (syn. Lonchidium Gerstaecker, 1854)

  • Kroyeria brasiliense Thatcher, 2006
  • Kroyeria carchariaeglauci Hesse, 1878
  • Kroyeria caseyi Benz & Deets, 1986
  • Kroyeria deetsi Dippenaar, Benz & Olivier, 2000
  • Kroyeria dispar Wilson C.B., 1935
  • Kroyeria echinata Rangnekar, 1956
  • Kroyeria elongata Pillai, 1967
  • Kroyeria gemursa Cressey, 1967
  • Kroyeria lineata Beneden, 1853
  • Kroyeria longicauda Cressey, 1970
  • Kroyeria minuta Pillai, 1968
  • Kroyeria papillipes Wilson C.B., 1932
  • Kroyeria spatulata Pearse, 1948
  • Kroyeria sphyrnae Rangnekar, 1957
  • Kroyeria sublineata Yamaguti & Yamasu, 1959

    PROKROYERIA Deets, 1987

  • Prokroyeria meridionalis (Ramirez, 1975)

    Krøyerina Wilson C.B., 1932 syn. Kroeyerina Wilson C.B., 1932; Lonchidium Gerstaecker, 1854 syn. Kroyeria Beneden, 1853.



    LERNAEOPODIDAE Milne Edwards, 1840 (syn. Naobranchiidae Yamaguti, 1939)


    MAXICLAVELLA Castro-Romero, Montes & Martorelli, 2022

  • Maxiclavella simplex (Castro-Romero & Baeza-Kuroki, 1985)

    PRAECLAVELLA Castro-Romero, Montes & Martorelli, 2022

  • Praeclavella bowmani (Kabata, 1963)
  • Praeclavella canaliculata (Wilson, C.B., 1915)
  • Praeclavella caudata (Castro-Romero et Baeza- Kuroki, 1985)
  • Praeclavella chiloensis (Castro-Romero, 1994)
  • Praeclavella convergentis (Castro-Romero, 1994)
  • Praeclavella embiotocae (Dojiri, 1981)
  • Praeclavella fortis (Castro et Gonzalez, 2009)
  • Praeclavella gracilis (Hansen, 1923)
  • Praeclavella hardingi (Nunes-Ruivo, 1964)
  • Praeclavella irina (Wilson, C.B., 1915)
  • Praeclavella levis (Wilson, C.B., 1915
  • Praeclavella longicauda (Ho, 1993)
  • Praeclavella nasalis Castro-Romero, Montes & Martorelli, 2022
  • Praeclavella ovata (Yamaguti, 1939)
  • Praeclavella parva (Wilson, C.B., 1915)
  • Praeclavella porogadi (Nunes-Ruivo, 1964)
  • Praeclavella scombropis (Yamaguti, 1939)
  • Praeclavella singularis (Castro-Romero et Gonzalez, 2009)
  • Praeclavella stichaei (Kroyer, 1863)
  • Praeclavella tumida (Wilson, C.B., 1915)

    ACESPADIA Leigh-Sharpe, 1933

  • Acespadia pomposa Leigh-Sharpe, 1933

    ALELLA Leigh-Sharpe, 1925

  • Alella berecynthia Leigh-Sharpe, 1936 (syn. Clavellomimus berecynthia (Leigh-Sharpe, 1936)
  • Alella canthari (Heller, 1865)
  • Alella gibbosa Olibvier 1995 [South Africa] (?)
  • Alella igillimpethu Erasmus et al., 2022 [South Africa]
  • Alella macrotrachelus (Brian, 1906)
  • Alella pagelli (Krøyer, 1863)
  • Alella tarakihi Hewitt & Blackwell, 1987 (syn. Clavellotis tarakihi (Hewitt & Blackwell, 1987)

    ANACLAVELLA Heegaard, 1940

  • Anaclavella filifera Kabata, 1992
  • Anaclavella sillaginoides Heegaard, 1940

    BRACHIELLA Cuvier, 1830

    (syn.Charopinopsis Yamaguti, 1963; Eobrachiella Ho & Do, 1984; Epibrachiella Wilson C.B., 1915; Thynnicola Miculicich, 1904

  • Brachiella albida (Rangnekar, 1956) (syn. Parabrachiella albida (Rangnekar, 1956)
  • Brachiella anisotremi (Castro-Romero & Baeza-Kuroki, 1989) (syn. Parabrachiella anisotremi (Castro-Romero & Baeza-Kuroki, 1989)
  • Brachiella annulata Markevich, 1940 (syn. Parabrachiella annulata (Markevich, 1940)
  • Brachiella anserina Wilson C.B., 1908 (syn. Parabrachiella anserina (Wilson C.B., 1908)
  • Brachiella concava Wilson (?)
  • Brachiella cybii Pillai, Prabha & Balaraman, 1982
  • Brachiella elegans Richiardi, 1880
  • Brachiella magna Kabata, 1968
  • Brachiella malayensis Ohtsuka, Piasecki, Ismail & Kamarudin, 2020
  • Brachiella neglecta Brian, 1898
  • Brachiella ovalis Beneden, 1870
  • Brachiella parva Nuñes-Ruivo, 1957
  • Brachiella quaternia (Wilson C.B., 1935)
  • Brachiella regia Lewis, 1967
  • Brachiella squali Wilson, 1944 (?)
  • Brachiella thynni Cuvier, 1830

    BRIANELLA Wilson C.B., 1915

  • Brianella corniger Wilson C.B., 1915
  • Brianella stebleri Monard, 1926 (syn. Brianola stebleri (Monard, 1926)

    CHAROPINUS Krøyer, 1863 (syn. Stylophorus Hesse, 1879; Thomsonella Wilson C.B., 1915)

  • Charopinus albidus Rangnekar, 1956 (syn. Brachiella albida (Rangnekar, 1956); Parabrachiella albida (Rangnekar, 1956)
  • Charopinus bicaudatus (Krøyer, 1837) (syn. Pseudocharopinus bicaudatus (Krøyer, 1837)
  • Charopinus cameroni Heller, 1949 (syn. Dendrapta cameroni (Heller, 1949)
  • Charopinus chimaerae Yamaguti, 1939 (syn. Schistobrachia chimaerae (Yamaguti, 1939)
  • Charopinus cluthae Hansen, 1923 (syn. Lernaeopodina longimana (Olsson, 1869)
  • Charopinus dalmanni (Retzius, 1830)
  • Charopinus dasyatis Pearse, 1952 [Australia] (?)
  • Charopinus dentatus (Wilson C.B., 1912) (syn. Pseudocharopinus dentatus (Wilson C.B., 1912)
  • Charopinus dubius Scott T., 1900
  • Charopinus malleus Rudolphi in Nordmann, 1832 (syn. Pseudocharopinus malleus (Rudolphi in Nordmann, 1832)
  • Charopinus markewitschi Gusev, 1951 (syn. Pseudocharopinus markewitschi (Gusev, 1951)
  • Charopinus narcinae Pillai, 1962 (syn. Pseudocharopinus narcinae (Pillai, 1962)
  • Charopinus parkeri (Thomson G.M., 1890)
  • Charopinus pastinacae (Beneden, 1851) (syn. Pseudocharopinus malleus (Rudolphi in Nordmann, 1832)
  • Charopinus quaternius Wilson C.B., 1935 (syn. Brachiella quaternia (Wilson C.B., 1935)
  • Charopinus ramosus Krøyer, 1863 (syn. Schistobrachia ramosa (Krøyer, 1863)

    CLAVELLA Oken, 1815 (syn. Anchorella Cuvier, 1830; Schisturus Oken, 1815)

    "The genus Clavella is one of the most complex and controversial groups among parasitic Copepoda. Approximately 116 species have been described in this group; however, over 50% of these species have been transferred to other genera. Thus, only 44 are considered valid species . Original descriptions of many species are poor, which have not been improved over time. In addition, incomplete descriptions have been applied for appendages; however, detailed descriptions are required to distinguish these species" (from:Munoz & Castro, 2022)

  • Clavella adunca (Strøm, 1762)
  • Clavella alata Brian, 1909
  • Clavella angulata (Krøyer, 1863)
  • Clavella appendiculosa (Krøyer, 1863)
  • Clavella appendiculata (Krøyer, 1863) (syn. Brachiella appendiculata (Krøyer, 1863); Neobrachiella appendiculata (Krøyer, 1863)
  • Clavella applicata Castro-Romero & Baeza-Kuroki, 1985
  • Clavella bathyalis Kazachenko & Avdeev, 1977
  • Clavella bergyltae (Krøyer, 1863)
  • Clavella bowmani Kabata, 1963
  • Clavella canaliculata Wilson C.B., 1915
  • Clavella canthari (Heller, 1865) (syn. Clavellodes canthari (Heller, 1865); Lernaeomyzon canthari (Heller, 1865); Alella canthari (Heller, 1865)
  • Clavella caudata Castro-Romero & Baeza-Kuroki, 1985
  • Clavella centrodonti (Beneden, 1870) (syn. Alella pagelli (Krøyer, 1863)
  • Clavella characis Brian, 1906 (syn. Clavellotis characis (Brian, 1906)
  • Clavella chiloensis Castro-Romero, 1994
  • Clavella cluthae Scott T., 1902 (syn. Hatschekia cluthae (Scott T., 1902)
  • Clavella collaris Ho, 1993
  • Clavella convergentis Castro-Romero, 1994
  • Clavella cornucola Munoz & Castro, 2021 [Chile]
  • Clavella cornuta (Müller O.F., 1776) (syn. Acanthochondria cornuta (Müller O.F., 1776)
  • Clavella delamarei Nuñes-Ruivo, 1953 (syn. Clavellomimus delamarei (Nuñes-Ruivo, 1953)
  • Clavella deminuta Kabata, 1992
  • Clavella denticis (Krøyer, 1863)
  • Clavella dilatata (Krøyer, 1863) (syn. Clavellotis dilatata (Krøyer, 1863)
  • Clavella diversia Ho, 1993
  • Clavella emarginata (Krøyer, 1837) (syn. Clavellisa emarginata (Krøyer, 1837)
  • Clavella embiotocae Dojiri, 1981
  • Clavella fallax (Heller, 1865) (syn. Clavellotis fallax (Heller, 1865)
  • Clavella fasiculata (Leidy, 1889) (syn. Brachiella fasiculata (Leidy, 1889)
  • Clavella fortis Castro & Gonzalez, 2009
  • Clavella gadomi Ho, 1993
  • Clavella gracilis Hansen, 1923
  • Clavella hardingi Nuñes-Ruivo, 1964
  • Clavella hippoglossi Cuvier, 1930 (syn. Hatschekia hippoglossi (Cuvier, 1830)
  • Clavella hostilis (Heller, 1865) (syn. Clavellopsis hostilis (Heller, 1865)
  • Clavella insolita Wilson C.B., 1915
  • Clavella intermedia (Quidor, 1906) (syn. Clavellomimus intermedia (Quidor, 1906)
  • Clavella irina Wilson C.B., 1915
  • Clavella inversa Wilson C.B., 1913 (syn. Mixtio inversa (Wilson C.B., 1913)
  • Clavella labracis Beneden, 1871 (syn. Hatschekia labracis (Beneden, 1871)
  • Clavella labrimixti (Hesse, 1878) (syn. Hatschekia labracis (Beneden, 1871)
  • Clavella laciniata (Krøyer, 1863) (syn. Clavellopsis laciniata (Krøyer, 1863)
  • Clavella levis Wilson C.B., 1915
  • Clavella lizae (Krøyer, 1863) (syn. Naobranchia lizae (Krøyer, 1863)
  • Clavella longicauda Ho, 1993
  • Clavella macrotrachelus Brian, 1906 (syn. Alella macrotrachelus (Brian, 1906)
  • Clavella macruri Hansen, 1923 (syn. Clavellomimus macruri (Hansen, 1923)
  • Clavella microcephala (Nordmann, 1832)
  • Clavella mulli van Beneden, 1851 (syn. Hatschekia mulli (van Beneden, 1851)
  • Clavella okamurai Ho, 1993
  • Clavella ovata Yamaguti, 1939
  • Clavella ovalis (Krøyer, 1837) (syn. Clavellisa emarginata (Krøyer, 1837)
  • Clavella pagelli (Krøyer, 1863) (syn. Alella pagelli (Krøyer, 1863)
  • Clavella pagri (Krøyer, 1863) (syn. Clavellotis pagri (Krøyer, 1863)
  • Clavella paradoxa (Beneden, 1851) (syn. Kabatazus paradoxus (van Beneden, 1851)
  • Clavella parva Wilson C.B., 1912
  • Clavella perfida Wilson C.B., 1915
  • Clavella pinguis Wilson C.B., 1915
  • Clavella porogadi Nuñes-Ruivo, 1964
  • Clavella robusta Wilson C.B., 1912 (syn. Parabrachiella robusta (Wilson C.B., 1912; syn. Neobrachiella robusta (Wilson 1912)
  • Clavella rugosa (Krøyer, 1837) (syn. Clavellodes rugosus (Krøyer, 1837)
  • Clavella sargi (Kurtz, 1877) (syn. Clavellotis sargi (Kurz, 1877)
  • Clavella scari Krøyer, 1838 (syn. Hatschekia scari (Krøyer, 1838)
  • Clavella sciaenae Brian, 1906 (syn.Parabrachiella sciaenae (Brian, 1906)
  • Clavella sciaenophila (Heller, 1865) (syn. Parabrachiella sciaenophila (Heller, 1865)
  • Clavella scombropis Yamaguti, 1939
  • Clavella scombri (Kurz, 1877) (syn. Clavellisa scombri (Kurz, 1877)
  • Clavella simplex Castro-Romero & Baeza-Kuroki, 1985
  • Clavella singularis Castro & Gonzalez, 2009
  • Clavella sokodara Ho, 1993
  • Clavella squamigera Wilson C.B., 1915
  • Clavella stellata (Krøyer, 1838)
  • Clavella stichaei (Krøyer, 1863)
  • Clavella strumosa Brian, 1906 (syn. Clavellotis strumosa (Brian, 1906)
  • Clavella tenuis Heller, 1865 (syn. Hatschekia tenuis (Heller, 1865)
  • Clavella triglae (Kurz, 1877) (syn. Parabrachiella triglae (Claus, 1860)
  • Clavella tumida Wilson C.B., 1915
  • Clavella tumidula Kabata, 1992
  • Clavella uncinata (Müller O.F., 1776)
  • Clavella urolophi (Krøyer, 1863)
  • Clavella zini Kabata, 1979

    CLAVELLISA Wilson C.B., 1915 (SYN. Epiclavella Yü, 1933)

  • Clavellisa chinensis (Yü, 1933)
  • Clavellisa cordata Wilson C.B., 1915
  • Clavellisa dorosomatis Yamaguti, 1939
  • Clavellisa dussumieriae Gnanamuthu, 1948
  • Clavellisa emarginata (Krøyer, 1837)
  • Clavellisa ilishae Pillai, 1962
  • Clavellisa obcordatus Rangnekar, 1957
  • Clavellisa pellone Tripathi, 1962
  • Clavellisa phasa Tripathi, 1962
  • Clavellisa pomolobi (Fowler, 1912)
  • Clavellisa scombri (Kurz, 1877)
  • Clavellisa spinosa Wilson C.B., 1915

    CLAVELLISTES Shiino, 1963 (syn. Andropoda Kabata, 1965 )

  • Clavellistes lampri (Scott T. & Scott A., 1901)
  • Clavellistes shoyoae Shiino, 1963

    CLAVELLODES Wilson C.B., 1915

  • Clavellodes canthari (Heller, 1865) (syn. Lernaeomyzon canthari (Heller, 1865)
  • Clavellodes ditrematis Yamaguti, 1939 (syn. Alella ditrematis (Yamaguti, 1939)
  • Clavellodes intermedia (Quidor, 1906) (syn. Clavellomimus intermedia (Quidor, 1906)
  • Clavellodes macrotrachelus (Brian, 1906) (syn. Alella macrotrachelus (Brian, 1906)
  • Clavellodes pagelli (Krøyer, 1863) (syn. Alella pagelli (Krøyer, 1863)
  • Clavellodes pterobrachiata Kabata, 1968 (syn. Alella pterobrachiata (Kabata, 1968)
  • Clavellodes rugosus (Krøyer, 1837)

    CLAVELLOMIMUS Kabata, 1969

  • Clavellomimus berecynthia (Leigh-Sharpe, 1936)
  • Clavellomimus delamarei (Nuñes-Ruivo, 1953)
  • Clavellomimus intermedia (Quidor, 1906)
  • Clavellomimus macruri (Hansen, 1923)

    CLAVELLOPSIS Wilson C.B., 1915 (syn. Isobranchia Heegaard, 1947)

  • Clavellopsis appendiculata Kirtisinghe, 1950 (syn. Isobranchia appendiculata Heegaard, 1947
  • Clavellopsis bilobata Pillai, 1962 (syn. Clavellotis bilobata (Pillai, 1962)
  • Clavellopsis branchiostegi Yamaguti, 1939 (syn. Clavellotis branchiostegi (Yamaguti, 1939)
  • Clavellopsis characis (Richiardi, 1880) (syn. Clavellotis characis (Brian, 1906)
  • Clavellopsis dasyaticus Rangnekar, 1957(syn. Pseudocharopinus dasyaticus (Rangnekar, 1957)
  • Clavellopsis fallax Heller, 1865 (syn. Clavellotis fallax (Heller, 1865)
  • Clavellopsis flexicervica Shiino, 1956
  • Clavellopsis flexicervica Shiino, 1956
  • Clavellopsis hostilis (Heller, 1865) (syn. Brachiella hostilis (Heller, 1865)
  • Clavellopsis hugu Yamaguti, 1939 (syn. Neobrachiella hugu (Yamaguti, 1939)
  • Clavellopsis johnii Yamaguti, 1939 (syn. Parabrachiella johnii (Yamaguti, 1939)
  • Clavellopsis longimana Bere, 1936
  • Clavellopsis nodula Do & Ho, 1983 (syn. Clavellotis dilatata (Krøyer, 1863)
  • Clavellopsis pagri (Krøyer, 1863) (syn. Clavellotis pagri (Krøyer, 1863)
  • Clavellopsis paradoxa (van Beneden, 1851) (syn. Kabatazus paradoxus (van Beneden, 1851)
  • Clavellopsis parasargi Roubal, 1981 (syn. Pseudomixtio parasargi (Roubal, 1981)
  • Clavellopsis parva (Wilson C.B., 1912) (syn. Clavella parva Wilson C.B., 1912
  • Clavellopsis pellucidula Shiino, 1956
  • Clavellopsis polymixiae Yamaguti, 1939
  • Clavellopsis producta Wilson C.B., 1915
  • Clavellopsis robusta (Wilson C.B., 1912) (syn. Parabrachiella robusta (Wilson C.B., 1912)
  • Clavellopsis saba Yamaguti, 1939
  • Clavellopsis sargi (Kurz, 1877) (syn. Clavellotis sargi (Kurz, 1877)
  • Clavellopsis robusta (Wilson C.B., 1912) (syn, Parabrachiella robusta (Wilson C.B., 1912)
  • Clavellopsis strumosa (Brian, 1906) (syn. Clavellotis strumosa (Brian, 1906)
  • Clavellopsis tomoi Stadler, 1973
  • Clavellopsis trichiuri Yamaguti, 1939 (syn. Parabrachiella trichiuri (Yamaguti, 1939)

    CLAVELLOTIS Castro-Romero & Baeza-Kuroki, 1984

  • Clavellotis bilobata (Pillai, 1962)
  • Clavellotis branchiostegi (Yamaguti, 1939)
  • Clavellotis briani Benmansour, Ben Hassine, Diebakate & Raibaut, 2001
  • Clavellotis characis (Brian, 1906)
  • Clavellotis dilatata (Krøyer, 1863)
  • Clavellotis fallax (Heller, 1865)
  • Clavellotis pagri (Krøyer, 1863)
  • Clavellotis sargi (Kurz, 1877)
  • Clavellotis sebastidis Castro & González, 2005
  • Clavellotis strumosa (Brian, 1906)
  • Clavellotis tarakihi (Hewitt & Blackwell, 1987)

    CRYPTOVA Kabata, 1992

  • Cryptova limbifera Kabata, 1992

    DENDRAPTA Kabata, 1964

  • Dendrapta cameroni (Heller, 1949)
  • Dendrapta longiclavata Kabata & Gusev, 1966
  • Dendrapta nasicola Irigoitia, Taglioretti & Timi, 2020

    EUBRACHIELLA Wilson C.B., 1915

  • Eubrachiella antarctica (Quidor, 1906)
  • Eubrachiella gaini gaini (Quidor, 1913)
  • Eubrachiella exigua (Brian, 1906) (syn. Parabrachiella exigua (Brian, 1906)
  • Eubrachiella exilis (Shiino, 1956) (syn. Parabrachiella exilis (Shiino, 1956)
  • Eubrachiella mugilis Kabata, Raibaut & Ben Hassine, 1971 (syn.Parabrachiella mugilis (Kabata, Raibaut & Ben Hassine, 1971)
  • Eubrachiella sublobulata Barnard, 1955 accepted as Parabrachiella sublobulata (Barnard, 1955)

    EUCLAVELLISA Heegaard, 1940

  • Euclavellisa australis Heegaard, 1940

    KABATAHOIA Kazachenko, 2001

  • Kabatahoia pectinata (Ho, 1985)

    KABATAZUS Özdikmen, 2008 (syn. Advena Kabata, 1979)

  • Kabatazus paradoxus (van Beneden, 1851)

    LERNAEOPODA Blainville, 1822 (syn. Lernaeopodella Heegaard, 1962)

  • Lernaeopoda alpina Olsson, 1877 (syn. Salmincola edwardsii (Olsson, 1869)
  • Lernaeopoda arcturi Miers, 1877 (syn. Salmincola edwardsii (Olsson, 1869)
  • Lernaeopoda beani Wilson C.B., 1908 (syn. Salmincola beani (Wilson C.B., 1908)
  • Lernaeopoda bicaudata Krøyer, 1837 (syn. Pseudocharopinus bicaudatus (Krøyer, 1837)
  • Lernaeopoda bicauliculata Wilson C.B., 1908 (syn. Salmincola bicauliculata (Wilson C.B., 1908)
  • Lernaeopoda bidiscalis Kane, 1892
  • Lernaeopoda bivia Leigh-Sharpe, 1930
  • Lernaeopoda brongniartii Blainville, 1822
  • Lernaeopoda californiensis Dana, 1852 (syn. Salmincola californiensis (Dana, 1852)
  • Lernaeopoda carpionis Krøyer, 1837 (syn. Salmincola carpionis (Krøyer, 1837)
  • Lernaeopoda centroscyllii Hansen, 1923 (syn. Neoalbionella centroscyllii (Hansen, 1923)
  • Lernaeopoda cluthae Scott T., 1900 (syn. Lernaeopodina longimana (Olsson, 1869)
  • Lernaeopoda coregoni Smith S.I., 1874 (syn. Salmincola extensa (Kessler, 1868)
  • Lernaeopoda coregonorum Kessler, 1868 (syn. Salmincola coregonorum (Kessler, 1868)
  • Lernaeopoda cyprinacea Hermann, 1783 (syn. Salmincola salmonea (Linnaeus, 1758)
  • Lernaeopoda dalmanni Retzius; of Krøyer, 1837 (syn. Charopinus dalmanni (Retzius, 1830)
  • Lernaeopoda edwardsii Olsson, 1869 (syn. Salmincola edwardsii (Olsson, 1869)
  • Lernaeopoda elongata (Grant, 1827) (syn. Ommatokoita elongata (Grant, 1827)
  • Lernaeopoda etmopteri Yamaguti, 1939 (syn. Neoalbionella etmopteri (Yamaguti, 1939)
  • Lernaeopoda extensa Kessler, 1868 (syn. Salmincola extensa (Kessler, 1868)
  • Lernaeopoda extumescens Gadd, 1901 (syn. Salmincola extumescens (Gadd, 1901)
  • Lernaeopoda falculata Wilson C.B., 1908 (syn. Salmincola californiensis (Dana, 1852)
  • Lernaeopoda fontinalis Smith S.I., 1874 (syn. Salmincola edwardsii (Olsson, 1869)
  • Lernaeopoda galei Krøyer, 1837
  • Lernaeopoda gibber Wilson C.B., 1908 (syn. Salmincola gibber (Wilson C.B., 1908)
  • Lernaeopoda globosa Leigh-Sharpe, 1918 (syn. Neoalbionella globosa (Leigh-Sharpe, 1918)
  • Lernaeopoda inermis Wilson C.B., 1911 (syn. Salmincola inermis (Wilson C.B., 1911)
  • Lernaeopoda lampri Scott T. & Scott A., 1901 (syn. Thysanote lampri (Scott T. & Scott A., 1901)
  • Lernaeopoda longibrachia Brian, 1912 ( syn. Lernaeopodina longibrachia (Brian, 1912)
  • Lernaeopoda longicaudata Hansen, 1923 (syn. Neoalbionella longicaudata (Hansen, 1923)
  • Lernaeopoda longimana Olsson, 1869 (syn. Lernaeopodina longimana (Olsson, 1869)
  • Lernaeopoda lotae Olsson, 1877 (syn. Salmincola lotae (Olsson, 1877)
  • Lernaeopoda major (Heegaard, 1962)
  • ernaeopoda maraenae Olsson, 1877 (syn. Salmincola extensa (Kessler, 1868)
  • Lernaeopoda musteli Thomson G.M., 1890
  • Lernaeopoda obesa Krøyer, 1837 (syn. Brachiella obesa (Krøyer, 1837)
  • Lernaeopoda oviformis Shiino, 1956 (syn. Neoalbionella oviformis (Shiino, 1956)
  • Lernaeopoda salmoneus (Gissler, 1751) (syn. Salmincola salmonea (Linnaeus, 1758)
  • Lernaeopoda scyllicola Leigh-Sharpe, 1916 (syn. Lernaeopoda galei Krøyer, 1837
  • Lernaeopoda similis Scott T. & Scott A., 1913 (syn. Lernaeopodina longimana (Olsson, 1869)
  • Lernaeopoda longibrachia (?)
  • Lernaeopoda siscowet Smith S.I., 1874 (syn. Salmincola siscowet (Smith S.I., 1874)
  • Lernaeopoda spinacis Brian, 1908 (syn. Lernaeopodina spinacis (Brian, 1908)
  • Lernaeopoda stromatei Gnanamuthu, 1950 (syn. Thysanote appendiculata (Steenstrup & Lütken, 1861)
  • Lernaeopoda tenuis Castro-Romero & Baeza-Kuroki, 1986 (syn. Neoalbionella tenuis (Castro-Romero & Baeza-Kuroki, 1986)
  • Lernaeopoda tuberculata Gadd, 1908 (syn. Salmincola nordmanni (Kessler, 1868)
  • Lernaeopoda upenaei Pillai, 1962

    Recently Hasegawa et al. (20229 evaluated Salmincola as a valid genus

    LERNAEOPODINA Wilson C.B., 1915

  • Lernaeopodina longibrachia (Brian, 1912)
  • Lernaeopodina longimana (Olsson, 1869)
  • Lernaeopodina pacifica Kabata & Gusev, 1966
  • Lernaeopodina pectinata Ho, 1985 (syn. Kabatahoia pectinata (Ho, 1985)
  • Lernaeopodina relata Wilson C.B., 1915

    Lernaeopodinae Wilson C.B., 1915

    MARGOLISIUS Benz, Kabata & Bullard, 2000

  • Margolisius abditus Benz, Kabata & Bullard, 2000

    MIXTIO Kabata, 1986

  • Mixtio inversa (Wilson C.B., 1913)

    NAOBRANCHIA Hesse, 1863 (syn. Axine Oken, 1815; Cestopoda Kurz, 1877)

  • Naobranchia alta Kabata, 1992
  • Naobranchia aulopi Yamaguti, 1939
  • Naobranchia auriculata Shiino, 1958
  • Naobranchia cygniformis Hesse, 1863
  • Naobranchia denticis Madinabeitia & Nagasawa, 2011
  • Naobranchia hemiconiati Nuñes-Ruivo, 1963
  • Naobranchia kabatana Dippenaar & Jordaan, 2008
  • Naobranchia lizae (Krøyer, 1863)
  • Naobranchia maxima Ho, 1975
  • Naobranchia microsoma Dojiri, 1981
  • Naobranchia occidentalis Wilson C.B., 1915
  • Naobranchia pagelli Nuñes-Ruivo, 1963
  • Naobranchia polynemi Tripathi, 1962
  • Naobranchia pomolobi Fowler, 1912 transferred to Clavellisa pomolobi (Fowler, 1912)
  • Naobranchia pritchardae Kensley & Grindley, 1973
  • Naobranchia sargi Nuñes-Ruivo, 1963
  • Naobranchia scorpaenae Dojiri, 1981
  • Naobranchia smaridis Nuñes-Ruivo, 1963
  • Naobranchia spinosa Pearse, 1952
  • Naobranchia variabilis Brian, 1924
  • Naobranchia vermiformis Rangnekar, 1956
  • Naobranchia wilsoni Nigrelli, 1933

    NECTOBRACHIA Fraser, 1920 (syn. Lernaeopodopsis Hansen, 1923)

  • Nectobrachia iceli Ho & Kim I.H., 1996
  • Nectobrachia indivisa Fraser, 1920
  • Nectobrachia producta (Hansen, 1923)

    NEOALBIONELLA Özdikmen, 2008 (syn. Albionella Kabata, 1979)

  • Neoalbionella benzipirata Ruiz & Bullard, in Ruiz, Driggers & Bullard, 2019
  • Neoalbionella centroscyllii (Hansen, 1923)
  • Neoalbionella dannytangi Ruiz & Bullard, 2019
  • Neoalbionella etmopteri (Yamaguti, 1939)
  • Neoalbionella fabricii (Rubec & Hogans, 1988)
  • Neoalbionella globosa (Leigh-Sharpe, 1918)
  • Neoalbionella izawai Dippenaar, 2020
  • Neoalbionella kabatai (Benz & Izawa, 1990)
  • Neoalbionella longicaudata (Hansen, 1923)
  • Neoalbionella oviformis (Shiino, 1956)
  • Neoalbionella tenuis (Castro-Romero & Baeza-Kuroki, 1986)

    NUDICLAVELLA Ho, 1975

  • Nudiclavella galapagoensis Ho, 1975

    OMMATOKOITA Leigh-Sharpe, 1926

  • Ommatokoita elongata (Grant, 1827)

    PARABRACHIELLA Wilson C.B., 1915 (syn. Neobrachiella Kabata, 1979; Probrachiella Wilson C.B., 1915)

  • Parabrachiella albida (Rangnekar, 1956)
  • Parabrachiella amphipacifica (Ho, 1982)
  • Parabrachiella anisotremi (Castro-Romero & Baeza-Kuroki, 1989)
  • Parabrachiella annulata (Markevich, 1940)
  • Parabrachiella appendiculata (Krøyer, 1863) (syn. Neobrachiella appendiculata (Krøyer, 1863)
  • Parabrachiella auriculata (Castro-Romero & Baeza-Kuroki, 1987)
  • Parabrachiella bera (Yamaguti, 1939)
  • Parabrachiella bispinosa (Nordmann, 1832)
  • Parabrachiella brevibrachiata (Kabata, 1992)
  • Parabrachiella brevicapita (Ho & Do, 1984)
  • Parabrachiella chavesii (Beneden, 1891)
  • Parabrachiella chevreuxii (Beneden, 1891)
  • Parabrachiella chlorophthalmi (Kabata, 1992)
  • Parabrachiella cirrata (Heegaard, 1962)
  • Parabrachiella cirrocauda (Heegaard, 1962)
  • Parabrachiella dentici (Nuñes-Ruivo, 1962)
  • Parabrachiella dispar (Castro-Romero & Baeza-Kuroki, 1987) (Neobrachiella dispar in thye oroginal description)
  • Parabrachiella elegans (Richiardi, 1880) (syn. Brachiella elegans Richiardi, 1880
  • Parabrachiella exigua (Brian, 1906)
  • Parabrachiella exilis (Shiino, 1956)
  • Parabrachiella fasciata (Castro-Romero & Baeza-Kuroki, 1987) (Neobraachiella fasciata in the original description)
  • Parabrachiella fasciculata (Leidy, 1889)
  • Parabrachiella gracilis (Wilson C.B., 1908)
  • Parabrachiella gulosa (Wilson C.B., 1915)
  • Parabrachiella gymnobrachiata (Kabata, 1968)
  • Parabrachiella hoi (Piasecki, 1993)
  • Parabrachiella hoplognathi (Yamaguti, 1939)
  • Parabrachiella hostilis (Heller, 1865)
  • Parabrachiella hugu (Yamaguti, 1939)
  • Parabrachiella impudica (Nordmann, 1832) (syn. Thysanote impudica (Nordmann, 1832)
  • Parabrachiella incurva Shiino, 1956
  • Parabrachiella indica (Tripathi, 1962)
  • Parabrachiella insidiosa (Heller, 1865)
  • Parabrachiella intermedia (Bere, 1936)
  • Parabrachiella jarai Piasecki, Mlynarczyk & Hayward, 2010
  • Parabrachiella johnii (Yamaguti, 1939)
  • Parabrachiella kabatai (Luque & Farfan, 1991)
  • Parabrachiella lagenformis (Szidat, 1955)
  • Parabrachiella lata (Song & Chen, 1976)
  • Parabrachiella lutiani (Pillai, 1985)
  • Parabrachiella malabarica Pillai, Prabha & Balaraman, 1982
  • Parabrachiella menticirrhi (Luque & Farfan, 1990)
  • Parabrachiella merluccii (Bassett-Smith, 1896)
  • Parabrachiella microdigitata (Kabata, 1992)
  • Parabrachiella mirifica (Kabata, 1992)
  • Parabrachiella mitrata (Wilson, C.B., 1915)
  • Parabrachiella multifimbriata (Bassett-Smith, 1896)
  • Parabrachiella nitida (Wilson C.B., 1915)
  • Parabrachiella oralis (Castro-Romero & Baeza-Kuroki, 1987) (Neobrachiella oralis in the original description)
  • Parabrachiella otolithi (Pillai, 1962)
  • Parabrachiella papillosa (Pearse, 1952) (syn. Thysanote papillosa (Pearse, 1952)
  • Parabrachiella paralichthyos (Castro-Romero & Baeza-Kuroki, 1986)
  • Parabrachiella pillaii (Kabata & Tareen, 1987)
  • Parabrachiella pinguis (Wilson, C.B., 1915)
  • Parabrachiella platensis Montes, Castro-Romero & Martorelli, 2017
  • Parabrachiella regia (Lewis, A.G., 1967)
  • Parabrachiella richiardii (Ben Hassine & Raibaut, 1978)
  • Parabrachiella robusta (Wilson C.B., 1912)
  • Parabrachiella rostrata (Krøyer, 1837)
  • Parabrachiella rotunda (Pearse, 1952)
  • Parabrachiella sciaenae (Brian, 1906)
  • Parabrachiella sciaenophila (Heller, 1865)
  • Parabrachiella septicauda (Shiino, 1956)
  • Parabrachiella seriolae (Yamaguti & Yamasu, 1960) (syn. Brachiella seriolae Yamaguti & Yamasu, 1960
  • Parabrachiella sihama Song & Chen, 1976
  • Parabrachiella spinicephala Ringuelet, 1945
  • Parabrachiella stellifera (Heegaard, 1962)
  • Parabrachiella sublobulata (Barnard, 1955)
  • Parabrachiella superba (Leigh-Sharpe, 1934)
  • Parabrachiella supplicans (Barnard, 1955)
  • Parabrachiella tetrici (Kabata, 1968)
  • Parabrachiella trichiuri (Yamaguti, 1939)
  • Parabrachiella triglae (Claus, 1860)
  • Parabrachiella upenaei (Pillai, 1962) (syn. Lernaeopoda upenaei Pillai, 1962
  • Parabrachiella yongxingensis (Song & Chen, 1976)

    PROCLAVELLODES Kabata, 1967

  • Proclavellodes pillai Kabata, 1967

    PSEUDOCHAROPINOIDES Castro-Romero & Baeza-Kuroki, 1987

  • Pseudocharopinoides myliobatides Castro-Romero & Baeza-Kuroki, 1987

    PSEUDOCHAROPINUS Kabata, 1964

  • Pseudocharopinus bicaudatus (Krøyer, 1837)
  • Pseudocharopinus concavus (Wilson C.B., 1913)
  • Pseudocharopinus dasyaticus (Rangnekar, 1957)
  • Pseudocharopinus dentatus (Wilson C.B., 1912)
  • Pseudocharopinus malleus (Rudolphi in Nordmann, 1832)
  • Pseudocharopinus markewitschi (Gusev, 1951)
  • Pseudocharopinus narcinae (Pillai, 1962)
  • Pseudocharopinus pillaii Kabata, 1979
  • Pseudocharopinus pteromylaei Raibaut & Essafi, 1979
  • Pseudocharopinus pteroplateae (Yamaguti & Yamasu, 1959)
  • Pseudocharopinus squali (Wilson C.B., 1944)

    PSEUDOLERNAEOPODA Castro-Romero & Baeza-Kuroki, 1986

  • Pseudolernaeopoda caudocapta Castro-Romero & Baeza-Kuroki, 1986

    PSEUDOLERNAEOPODINA Hogans, 1988

  • Pseudolernaeopodina synophobranchi Hogans, 1988

    PSEUDOMIXTIO Kabata, 1990

  • Pseudomixtio parasargi (Roubal, 1981)

    SALMINCOLA Wilson C.B., 1915 (syn. Acanthochondria tiddi (Price & Hames, 1968); Entomoda tiddi Price & Hames, 1968; Pediculus Gissler, 1751)

  • Salmincola californiensis (Dana, 1852)
  • Salmincola carpionis (Krøyer, 1837)
  • Salmincola edwardsii (Olsson, 1869)
  • Salmincola extensa (Kessler, 1868)
  • Salmincola extumescens (Gadd, 1901)
  • Salmincola gordoni Gurney, 1933
  • Salmincola lavaretus Burdukovskaya & Pronin, 2010
  • Salmincola cf. markewitschi Shedko et Shedko, 2002
  • Salmincola mica Shedko, 2004
  • Salmincola salmonea (Linnaeus, 1758)
  • Salmincola svetlanovi Burdukovskaya & Pronin, 2010

    SCHISTOBRACHIA Kabata, 1964

  • Schistobrachia chimaerae (Yamaguti, 1939)
  • Schistobrachia jordaanae Dippenaar, Olivier & Benz, 2004
  • Schistobrachia kabata Dippenaar, 2016
  • Schistobrachia pilgrimi Kabata, 1988
  • Schistobrachia ramosa (Krøyer, 1863)
  • Schistobrachia tertia Kabata, 1970

    SPARIDICOLA Kabata & Tareen, 1987

  • Sparidicola lithognathi (Kensley & Grindley, 1973)

    THYSANOTE Krøyer, 1863 (syn. Brachiellina Pearse, 1952 ; Thysanotella Wilson C.B., 1915)

  • Thysanote alternans Kabata & Tareen, 1981
  • Thysanote appendiculata (Steenstrup & Lütken, 1861)
  • Thysanote chalermwati Piasecki, Ohtsuka & Yoshizaki, 2008
  • Thysanote eleutheronema Rangnekar, 1961
  • Thysanote epinepheli Yamaguti, 1939
  • Thysanote exornata Brian, 1939
  • Thysanote fimbriata (Heller, 1865)
  • Thysanote furcata Kirtisinghe, 1964
  • Thysanote gymnobrachiata Kabata, 1968 (syn. Parabrachiella gymnobrachiata (Kabata, 1968)
  • Thysanote goodi Causey, 1960
  • Thysanote heterodactyla Kirtisinghe, 1964
  • Thysanote impudica (Nordmann, 1832)
  • Thysanote lampri (Scott T. & Scott A., 1901) (syn. Clavellistes lampri (Scott T. & Scott A., 1901)
  • Thysanote lobiventris (Heller, 1865)
  • Thysanote longimana Wilson C.B., 1913
  • Thysanote magna (Song & Chen, 1976)
  • Thysanote multifimbriata (Bassett-Smith, 1898) (syn. Parabrachiella multifimbriata (Bassett-Smith, 1896)
  • Thysanote nudibranchiata Pillai, Prabha & Balaraman, 1982
  • Thysanote papillosa (Pearse, 1952)
  • Thysanote polyfimbriata Prabha & Pillai, 1981
  • Thysanote pomacanthi Krøyer, 1863
  • Thysanote ramosa (Richiardi, 1880) (syn. Neobrachiella ramosa (Richiardi, 1880)
  • Thysanote rastrelligeri Rangnekar, 1961
  • Thysanote triloba Pearse, 1952

    Tracheliastinae Wilson C.B., 1915

    VANBENEDENIA Malm, 1861

  • Vanbenedenia chimaerae (Heegaard, 1962)
  • Vanbenedenia grandis (Wilson C.B., 1915)
  • Vanbenedenia hydrolagae Oldewage, 1993
  • Vanbenedenia kroeyeri Malm, 1861


    Advena Kabata, 1979 syn. Kabatazus Özdikmen, 2008; Albionella Kabata, 1979 syn. Neoalbionella Özdikmen, 2008; Anchorella Cuvier, 1830 syn. Clavella Oken, 1815; Andropoda Kabata, 1965 syn.Clavellistes Shiino, 1963; Axine Oken, 1815 syn. Naobranchia Hesse, 1863; Brachiellina Pearse, 1952 syn.Thysanote Krøyer, 1863; Cestopoda Kurz, 1877 syn, Naobranchia Hesse, 1863; Charopinopsis Yamaguti, 1963 syn. Brachiella Cuvier, 1830; Eobrachiella Ho & Do, 1984 syn. Brachiella Cuvier, 1830; Epibrachiella Wilson C.B., 1915 syn. Brachiella Cuvier, 1830; Epiclavella Yü, 1933 syn. Clavellisa Wilson C.B., 1915 ; Isobranchia Heegaard, 1947 syn. Clavellopsis Wilson C.B., 1915; Lernaeopodella Heegaard, 1962 syn. Lernaeopoda Blainville, 1822; Lernaeopodopsis Hansen, 1923 syn. Nectobrachia Fraser, 1920 ; Neobrachiella Kabata, 1979 syn. Parabrachiella Wilson C.B., 1915; Pediculus Gissler, 1751 syn. Salmincola Wilson C.B., 1915; Probrachiella Wilson C.B., 1915 syn. Parabrachiella Wilson C.B., 1915; Stylophorus Hesse, 1879 syn. Charopinus Krøyer, 1863: Thomsonella Wilson C.B., 1915 syn.Charopinus Krøyer, 1863 Thynnicola Miculicich, 1904 syn. Brachiella Cuvier, 1830; Thysanotella Wilson C.B., 1915 syn. Thysanote Krøyer, 1863


    TRACHELIASTES Nordmann, 1832

  • Tracheliastes brevicorpus Kuang, 1980
  • Tracheliastes chimaerae Heegaard, 1962 accepted as Vanbenedenia chimaerae (Heegaard, 1962)
  • Tracheliastes chondrostomi Hanek, 1969
  • Tracheliastes fecundus Odenwall, 1931 accepted as Tracheliastes maculatus Kollar, 1835
  • Tracheliastes gigas Richiardi, 1881 accepted as Pseudotracheliastes stellatus (Mayor, 1824)
  • Tracheliastes grandis Wilson C.B., 1915 accepted as Vanbenedenia grandis (Wilson C.B., 1915)
  • Tracheliastes longicollis Markevich, 1940
  • Tracheliastes maculatus Kollar, 1835
  • Tracheliastes mourkii Hoffman, 1881
  • Tracheliastes polycolpus Nordmann, 1832
  • Tracheliastes sachalinensis Markevich, 1936
  • Tracheliastes soldatovi Markevich, 1933
  • Tracheliastes stellatus (Mayor, 1824) accepted as Pseudotracheliastes stellatus (Mayor, 1824)
  • Tracheliastes stellifer Kollar, 1835 accepted as Pseudotracheliastes stellifer (Kollar, 1835)
  • Tracheliastes tibetanus Kuang, 1964

    Acespadiinae Kazachenko, 2001 (Subfamily not accepted by Boxshall & Halsey, 2004)

  • Advena Kabata, 1979 accepted as Kabatazus Özdikmen, 2008
  • Albionella Kabata, 1979 accepted as Neoalbionella Özdikmen, 2008
  • Anchorella Cuvier, 1830 accepted as Clavella Oken, 1815
  • Andropoda Kabata, 1965 accepted as Clavellistes Shiino, 1963
  • Axine Oken, 1815 accepted as Naobranchia Hesse, 1863
  • Brachiellina Pearse, 1952 accepted as Thysanote Krøyer, 1863

    Brachiellinae Kazachenko, 2001 (Subfamily not accepted by Boxshall & Halsey, 2004)

  • Cestopoda Kurz, 1877 accepted as Naobranchia Hesse, 1863

    Charopininae Kazachenko, 2001 (Subfamily not accepted by Boxshall & Halsey, 2004)

  • Charopinopsis Yamaguti, 1963 accepted as Brachiella Cuvier, 1830

    Clavellisinae Kazachenko, 2001 (Subfamily not accepted by Boxshall & Halsey, 2004)

    Cryptovinae Kazachenko, 2001 (Subfamily not accepted by Boxshall & Halsey, 2004)

  • Eobrachiella Ho & Do, 1984 accepted as Brachiella Cuvier, 1830
  • Epibrachiella Wilson C.B., 1915 accepted as Brachiella Cuvier, 1830
  • Epiclavella Yü, 1933 accepted as Clavellisa Wilson C.B., 1915
  • Isobranchia Heegaard, 1947 accepted as Clavellopsis Wilson C.B., 1915

    Kabatahoinae Kazachenko, 2001 (Subfamily not accepted by Boxshall & Halsey, 2004)
  • Lernaeopodella Heegaard, 1962 accepted as Lernaeopoda Blainville, 1822
  • Lernaeopodopsis Hansen, 1923 accepted as Nectobrachia Fraser, 1920

    Nectobrachiinae Markevich, 1956 accepted as Lernaeopodinae Wilson C.B., 1915 (Subfamily not accepted by Yamaguti, 1963)

    Neobrachiella Kabata, 1979 accepted as Parabrachiella Wilson C.B., 1915

  • Pediculus Gissler, 1751 accepted as Salmincola Wilson C.B., 1915
  • Probrachiella Wilson C.B., 1915 accepted as Parabrachiella Wilson C.B., 1915

    Salmincolinae Kazachenko, 2001 (Subfamily not accepted by Boxshall & Halsey, 2004)

    Schistobrachiinae Kazachenko, 2001 (Subfamily not accepted by Boxshall & Halsey, 2004)

  • Schisturus Oken, 1815 accepted as Clavella Oken, 1815
  • Stylophorus Hesse, 1879 accepted as Charopinus Krøyer, 1863
  • Thomsonella Wilson C.B., 1915 accepted as Charopinus Krøyer, 1863
  • Thynnicola Miculicich, 1904 accepted as Brachiella Cuvier, 1830
  • Thysanotella Wilson C.B., 1915 accepted as Thysanote Krøyer, 1863

    Vanbenedeniinae Kazachenko, 2001 (Subfamily not accepted by Boxshall & Halsey, 2004)



    LERNANTHROPIDAE Kabata, 1979

    "The family Lernanthropidae Kabata, 1979 currently comprises about 150 species belonging to eight genera (Boxshall & Halsey, 2004). The genus Lernanthropus de Blainville, 1822 is the largest of the family, containing approximately 120 valid species, all of which parasitise marine fish hosts belonging to at least 31 different teleost families (Boxshall & Halsey, 2004)" (from: El Rashdi & Boxshall, 2016)

    AETHON Krøyer, 1837

  • Aethon bicamera Boxshall, Bernot, Barton, Diggles, Yong, Atkinson-Coyle & Hutson, 2020
  • Aethon garricki Hewitt, 1968
  • Aethon globosus (Pillai, 1967)
  • Aethon morelandi Hewitt, 1968
  • Aethon percis (Thomson G.M., 1890)
  • Aethon percis (Thomson G.M., 1890)
  • Aethon quadratus Krøyer, 1837

    LERNANTHROPINUS Ho & Do, 1985

  • Lernanthropinus carangoides Izawa, 2014
  • Lernanthropinus caranxi (Hesse, 1878)
  • Lernanthropinus decapteri (Pillai, 1964)
  • Lernanthropinus forficatus (Redkar, Rangnekar & Murti, 1949)
  • Lernanthropinus holiulini Izawa, 2018
  • Lernanthropinus labracoglossae Izawa, 2014
  • Lernanthropinus maajii Izawa, 2018
  • Lernanthropinus nematistii Deets & Benz, 1988
  • Lernanthropinus sphyraenae (Yamaguti & Yamasu, 1959)
  • Lernanthropinus temminckii (von Nordmann, 1864)
  • Lernanthropinus trachuri (Brian, 1903)

    LERNANTHROPODES Bere, 1936 (syn. Chauvanium Kazachenko, Kovaleva, Nguyen & Ngo, 2017)

  • Lernanthropodes chauvani (Kazachenko, Kovaleva, Nguyen & Ngo, 2017)
  • Lernanthropodes chorinemi Pillai, 1962
  • Lernanthropodes cucullus Bere, 1936
  • Lernanthropodes natalensis Kensley & Grindley, 1973
  • Lernanthropodes trachinoti Pillai, 1962

    LERNANTHROPSIS Ho & Do, 1985

  • Lernanthropsis mugilii (Shishido, 1898)

    LERNANTHROPUS de Blainville, 1822 (syn. Epachthes Nordmann, 1832 (synonym) Sanya Song & Chen, 1976 )

  • Lernanthropus abitocephalus Tripathi, 1962
  • Lernanthropus alatus Pillai, 1964
  • Lernanthropus alepicolus Boxshall, Bernot, Barton, Diggles, Yong, Atkinson-Coyle & Hutson, 2020
  • Lernanthropus amplitergum Pearse, 1951 [Bahamas]
  • Lernanthropus angulatus Krøyer, 1863 (syn. Sagum angulatum (Krøyer, 1863)
  • Lernanthropus antofagastiensis Castro-Romero & Baeza-Kuroki, 1985
  • Lernanthropus atrox Heller, 1865 [
  • Lernanthropus barnardi Capart, 1959
  • Lernanthropus belones Krøyer, 1863
  • Lernanthropus bifidus Pearse, 1951 [Bahamas]
  • Lernanthropus brevicephalus Rangnekar, 1957
  • Lernanthropus brevicornutus Kabata, 1979
  • Lernanthropus brevoortiae Rathburn (?)
  • Lernanthropus breviculus Kabata, 1979
  • Lernanthropus brevis Richiardi, 1879
  • Lernanthropus brevoortiae Rathbun, 1887
  • Lernanthropus cadenati Delamare Deboutteville & Nunes-Ruvio, 1954 [
  • Lernanthropus caesionis Yamaguti, 1954 (syn. Sagum caesionis (Yamaguti, 1954)
  • Lernanthropus callionymicola El-Rashidy & Boxshall, 2012
  • Lernanthropus capistroides Olivier & van Niekerk, 1995
  • Lernanthropus caudatus Wilson C.B., 1922
  • Lernanthropus chacchi Suárez-Morales, Reyes-Lizama & González-Solís, 2010
  • Lernanthropus chirocentrosus Tripathi, 1962
  • Lernanthropus chrysophrys Shishido, 1898
  • Lernanthropus corniger Yamaguti, 1954
  • Lernanthropus corteziensis Deets & Kabata, 1991
  • Lernanthropus cruciatus Pillai, 1962
  • Lernanthropus cynoscicola Timi & Etchegoin, 1996
  • Lernanthropus delamari Capart, 1959
  • Lernanthropus dussumieria Gnanamuthu, 1949
  • Lernanthropus ecclesi Kenseley & Grindley, 1972 [South Africa] (?)
  • Lernanthropus eddiwarneri Delamare Deboutteville & Nunes-Ruivo, 1954
  • Lernanthropus elegans Boxshall, Bernot, Barton, Diggles, Yong, Atkinson-Coyle & Hutson, 2020 [
  • Lernanthropus equulus (Song & Chen, 1976)
  • Lernanthropus engraulis Tripathi, 1962 (syn. Mitrapus engraulis (Tripathi, 1962)
  • Lernanthropus erythrocles Izawa, 2014
  • Lernanthropus francai Nuñes-Ruivo, 1962
  • Lernanthropus frondeus Wilson C.B., 1913
  • Lernanthropus gazzis Song & Chen, 1976
  • Lernanthropus giganteus Krøyer, 1863
  • Lernanthropus gisleri Beneden, 1852 [
  • Lernanthropus gnathanodontus Boxshall, Bernot, Barton, Diggles, Yong, Atkinson-Coyle & Hutson, 2020 [
  • Lernanthropus grassei Delamare Deboutteville, 1954
  • Lernanthropus guacoldae Villalba & Fernandez, 1984 [Chile] (?) [
  • Lernanthropus heteropodus Yü, 1933 (syn. Mitrapus heteropodus (Yü, 1933)
  • Lernanthropus hiatus Pearse, 1951 [Bahamas]
  • Lernanthropus hirsutus Pearse, 1952
  • Lernanthropus holmbergii Nordmann, 1864
  • Lernanthropus huamani Luque & Farfan, 1990
  • Lernanthropus ilishae Chin, 1948
  • Lernanthropus incilis Ho, Liu & Lin, 2011
  • Lernanthropus indefinitus Koyuncu, Castro Romero & Karaytug, 2012
  • Lernanthropus indicus Pillai, 1967
  • Lernanthropus koenigii Steenstrup & Lütken, 1861
  • Lernanthropus kroyeri Beneden, 1851
  • Lernanthropus larvatus Heller, 1865
  • Lernanthropus latis Yamaguti, 1954 [
  • Lernanthropus lativentris Heller, 1865 (syn. Sagum lativentris (Heller, 1865)
  • Lernanthropus leidyi Wilson C.B., 1922
  • Lernanthropus leiognathi Tripathi, 1962
  • Lernanthropus longilamina Pearse, 1951 [Bahamas]
  • Lernanthropus longipes Wilson C.B., 1932
  • Lernanthropus manicatus Wilson C.B., 1935
  • Lernanthropus microlamini Hewitt, 1968
  • Lernanthropus micropterygis Richiardi, 1884
  • Lernanthropus mitratus Shiino, 1959
  • Lernanthropus mollis Kabata, 1979
  • Lernanthropus monodi Delamare Deboutteville & Nunes-Ruivo, 1954
  • Lernanthropus musca Blainville, 1822
  • Lernanthropus nemipteri Jayasree & Pillai, 1976
  • Lernanthropus nobilis Heller, 1865
  • Lernanthropus nordmanni Wilson C.B., 1922
  • Lernanthropus nunesi Capart, 1959
  • Lernanthropus oblongus Pillai, 1964 (syn. Mitrapus oblongus (Pillai, 1964)
  • Lernanthropus obscurus Wilson C.B., 1913
  • Lernanthropus opisthopteri Pillai, 1964
  • Lernanthropus otolithi Pillai, 1963
  • Lernanthropus pacificus Oliva & Duran, 1982
  • Lernanthropus pagelli Krøyer, 1863
  • Lernanthropus pagodus Krøyer, 1863
  • Lernanthropus pami Tripathi, 1962
  • Lernanthropus paracruciatus Boxshall, Bernot, Barton, Diggles, Yong, Atkinson-Coyle & Hutson, 2020
  • Lernanthropus paradoxus (von Nordmann, 1832)
  • Lernanthropus paralonchuri Luque, Bruno & Covarrubia, 1989
  • Lernanthropus pemphericola Boxshall, Bernot, Barton, Diggles, Yong, Atkinson-Coyle & Hutson, 2020
  • Lernanthropus percis Thomson G.M., 1890 (syn.Aethon percis (Thomson G.M., 1890)
  • Lernanthropus petersi Beneden, 1857 (syn. Sagum petersi (Beneden, 1857)
  • Lernanthropus pillaii Kabata, 2005
  • Lernanthropus polynemi Richiardi, 1881
  • Lernanthropus pomadasysis Rangnekar & Murti, 1961
  • Lernanthropus pomatomi Rathbun, 1887
  • Lernanthropus priacanthi Kirtisinghe, 1956 (syn. Norion priacanthi (Kirtisinghe, 1956)
  • Lernanthropus pristipomoides Kirtisinghe, 1937
  • Lernanthropus pupa Burmeister, 1833
  • Lernanthropus quadratus Krøyer, 1837 (syn. Aethon quadratus Krøyer, 1837
  • Lernanthropus rathbuni Wilson C.B., 1922
  • Lernanthropus robustus Pillai, 1964
  • Lernanthropus rubiginosus Redkar, Rangnekar & Murti, 1949 (syn. Mitrapus rubiginosus (Redkar, Rangnekar & Murti, 1949)
  • Lernanthropus sanguineus Song, in Song & Chen, 1976 (syn. Sagum sanguineus (Song, in Song & Chen, 1967)
  • Lernanthropus sarbae Kensley & Grindley, 1973
  • Lernanthropus sciaeni Gnanamuthu, 1947
  • Lernanthropus scribae Krøyer, 1863
  • Lernanthropus secutoris Pillai, 1963
  • Lernanthropus selenotoca Boxshall, Bernot, Barton, Diggles, Yong, Atkinson-Coyle & Hutson, 2020
  • Lernanthropus senegalensis Diebakate & Raibaut, 1996
  • Lernanthropus seriolii Shishido, 1898
  • Lernanthropus sillaginis Pillai, 1963
  • Lernanthropus sphyraenae Chin, 1948
  • Lernanthropus spiculatus Wilson C.B., 1913
  • Lernanthropus talipes Wilson C.B., 1935
  • Lernanthropus tenuis Wilson C.B., 1922
  • Lernanthropus theodori Brian, 1924
  • Lernanthropus togatus Kabata, 1971
  • Lernanthropus toguebayei Diebakate & Raibaut, 1996
  • Lernanthropus triangularis Pillai, 1963
  • Lernanthropus tylosuri Richiardi, in Goggio, 1906
  • Lernanthropus villiersi Delamare Deboutteville & Nunes-Ruivo, 1954
  • Lernanthropus wilsoni Pearse, 1952

    MITRAPUS Song & Chen, 1976

  • Mitrapus engraulis (Tripathi, 1962)
  • Mitrapus heteropodus (Yü, 1933)
  • Mitrapus oblongus (Pillai, 1964)

    NORION Nordmann, 1864

  • Norion expansus Nordmann, 1864
  • Norion globosus Pillai, 1967 (syn. Aethon globosus (Pillai, 1967)
  • Norion priacanthi (Kirtisinghe, 1956) (syn. Aethon priacanthi (Kirtisinghe, 1956)
  • Norion tayenus Ho & Kim I.H., 2004

    SAGUM Wilson C.B., 1913 (syn. Paralernanthropus Hewitt, 1968; Parlernanthropus Hewitt, 1968; Pseudolernanthropus Yamaguti & Yamasu, 1960; Stalagmus Nordmann, 1864

  • Sagum angulatum (Krøyer, 1863)
  • Sagum bitaro Uyeno & Naruse, 2018
  • Sagum brotulae Izawa, 2018
  • Sagum caesionis (Yamaguti, 1954)
  • Sagum enneacentri Pillai, 1985
  • Sagum epinepheli (Yamaguti & Yamasu, 1960)
  • Sagum flagellatum Wilson C.B., 1913
  • Sagum foliaceum (Goggio, 1906)
  • Sagum folium Ho, Liu & Lin, 2011
  • Sagum gurukun Uyeno & Naruse, 2018
  • Sagum lativentris (Heller, 1865)
  • Sagum paracaesionis Izawa, 2014
  • Sagum petersi (Beneden, 1857)
  • Sagum pillaisebastiani Izawa, 2018
  • Sagum poeyi Ortiz et al., 2003 [Cuba] (?)
  • Sagum posteli Delamare Deboutteville & Nunes-Ruivo, 1954
  • Sagum sanguineus (Song, in Song & Chen, 1967)
  • Sagum texanum Pearse, 1952
  • Sagum vespertilio Kabata, 1979
  • Sagum vietnamiensis Kazachenko, Kovaleva, Nguyen & Ngo, 2017

    Key to the species of Sagum (adult females)


    Genus Chauvanium Kazachenko, Kovaleva, Nguyen & Ngo, 2017 syn. Bere, 1936; Genus Epachthes Nordmann, 1832 syn. Lernanthropus de Blainville, 1822; Genus Paralernanthropus Hewitt, 1968 siun. Sagum Wilson C.B., 1913 ; Genus Parlernanthropus Hewitt, 1968 syn. Paralernanthropus Hewitt; Genus Pseudolernanthropus Yamaguti & Yamasu, 1960 Sagum syn. Wilson C.B., 1913; Genus Sanya Song & Chen, 1976 syn. Lernanthropus de Blainville, 1822.


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