MEGAPONTIIDAE Heptner 1968
HYALOPONTIUS Sars G.O., 1909 (syn. Megapontius Hulsemann, 1965)
Hyalopontius alatus Boxshall, 1979
Hyalopontius boxshalli Humes, 1988 
Hyalopontius cinctus Boxshall, 1979
Hyalopontius enormis Boxshall, 1979
Hyalopontius hulsemannae Boxshall, 1979
Hyalopontius pleurospinosus (Heptner, 1968)
Hyalopontius roei Boxshall, 1979
Hyalopontius spinatus Boxshall, 1979
Hyalopontius typicus Sars G.O., 1909
MICROPONTIIDAE Gooding, 1957
MICROPONTIUS Gooding, 1957
Micropontius glaber Stock, Humes & Gooding, 1963
Micropontius ovoides Gooding, 1957
NANASPIDIDAE Humes & Cressey, 1959
ALLANTOGYNUS Changeux, 1960
Allantogynus delamarei Changeux, 1960
HONSHIA Avdeev,
Honshia lobata Avdeev, 2017
HUMESIA Avdeev, 1980
Humesia alata Avdeev, 1980
Humesia bicuspidata Avdeev, 1980
NANASPIS Humes & Cressey, 1959
Nanaspis boholensis Humes, 1980
Nanaspis exigua Stock, Humes & Gooding, 1962
Nanaspis manca Humes, 1973
Nanaspis media Stock, Humes & Gooding, 1962
Nanaspis mediterranea Stock & Kleeton, 1963
Nanaspis mixta Humes, 1975
Nanaspis moluccana Humes, 1980
Nanaspis ninae Bresciani & Lützen, 1962
Nanaspis pollens Stock, Humes & Gooding, 1962
Nanaspis pusilla Humes, 1973
Nanaspis spinifera Humes, 1973
Nanaspis tonsa Humes & Cressey, 1959
Nanaspis truncata Stock, Humes & Gooding, 1962
KEY TO SPECIES OF NANASPIS
STEPHOPONTIUS Thompson I. & Scott A., 1903
Stephopontius ahni Kim I.H., 2010
Stephopontius typicus Thompson I.C. & Scott A., 1903
NICOTHOIDAE Dana, 1852 (syn. Choniostomatidae Hansen, 1886)Copepods in the family Nicothoidae Dana, 1849 are parasitic, and are classified in 22 genera
(Boxshall & Halsey, 2004; Ohtsuka et al., 2005; Bamber & Boxshall, 2006; Boyko, 2009).
ARHIZORHINA Bamber & Boxshall, 2006
Arhizorhina mekonicola Bamber & Boxshall, 2006
ASPIDOECIA Giard & Bonnier, 1889
Aspidoecia normani Giard & Bonnier, 1889
CEPHALORHIZA Boxshall & Harrison, 1988
Cephalorhiza flaccida Boxshall & Harrison, 1988
CHONOMYZON Pillai, 1962
Choniomyzon inflatus Wakabayashi, Otake, Tanaka & Nagasawa, 2013
Choniomyzon libiniae Santos & Björnberg, 2004
Choniomyzon panuliri Pillai, 1962
Choniomyzon taiwanensis Cheng, Wakabayashi & Pan, 2021
CHONIORHIZA Boxshall & Lincoln, 1983
Choniorhiza enaulis Boxshall & Lincoln, 1983
CHONIOSPHAERA Connolly, 1929 (Lecithomyzon Bloch & Gallien, 1933)
Choniosphaera cancrorum Connolly, 1929
Choniosphaera indica Gnanamuthu, 1954
Choniosphaera maenadis (Bloch & Gallien, 1933)
CHONIOSTOMA Hansen, 1886 (syn. Heptalobus Nierstrasz & Brender ŕ Brandis, 1930)
Choniostoma hanseni Giard & Bonnier, 1889
Choniostoma mirabile Hansen, 1886
Choniostoma paradoxum (Nierstrasz & Brender ŕ Brandis, 1930)
Choniostoma rotundatum Stock, 1958
DIEXANTHEMA Ritchie, 1975
Diexanthema apoda Boxshall & Harrison, 1988
Diexanthema bathydiaita Ritchie, 1975
Diexanthema corrugatum Boxshall & Harrison, 1988
Diexanthema desistoma Ritchie, 1975
Diexanthema hokuhomaruae Kalem et al., 2023 [Nort West Pacific]
Diexanthema nudum Boxshall & Harrison, 1988
Diexanthema ritchiei Boxshall & Harrison, 1988
HADROTHOE Humes, 1975
Hadrothoe crenulata Boxshall & Defaye, 1995
Hadrothoe crosnieri Humes, 1975
HANSENULUS Heron & Damkaer, 1986
Hansenulus trebax Heron & Damkaer, 1986
HOMOEOSCELIS Hansen, 1897
Homoeoscelis elongata Boxshall & Defaye, 1995
Homoeoscelis frigida Hansen, 1923
Homoeoscelis longipes Hansen, 1923
Homoeoscelis mediterranea Hansen, 1897
Homoeoscelis meridionalis Alberico & Roccatagliata, 2010
Homoeoscelis minuta Hansen, 1897
Homoeoscelis sedentaria (Bonnier, 1898)
MYSIDION Hansen, 1897
Mysidion abyssorum Hansen, 1897
Mysidion commune Hansen, 1897
NEOMYSIDION Ohtsuka, Boxshall & Harada, 2005
Neomysidion rahotsu Ohtsuka, Boxshall & Harada, 2005
NICORHIZA Lincoln & Boxshall, 1983
Nicorhiza humesi Lincoln & Boxshall, 1983
Nicorhiza humesi Lincoln & Boxshall, 1983
NICOTHOE Audouin & Milne Edwards H., 1826
Nicothoe analata Kabata, 1966
Nicothoe astaci Audouin & Edwards, 1826
Nicothoe brucei Kabata, 1967
Nicothoe procircularis Carton, 1967 (syn. Pseudonicothoe procircularis (Carton, 1967)
Nicothoe simplex Kabata, 1967
Nicothoe tumulosa Cressey, 1976
PARANICOTHOE Carton, 1970 (syn. Pseudonicothoe Avdeev & Avdeev, 1978)
Paranicothoe branchialis (Avdeev & Avdeev, 1978) (syn. Pseudonicothoe branchialis Humes & Boxshal, 1996)
Paranicothoe procircularis (Carton, 1967)
RHIZORHINA Hansen, 1892 (Salenskya Giard & Bonnier, 1893)
Rhizorhina aesthetes Boxshall & Harrison, 1988
Rhizorhina ampeliscae Hansen, 1892
Rhizorhina hystrix Boxshall & Harrison, 1988
Rhizorhina leptostracae Gotto, 1984
Rhizorhina ohtsukai Kakui, 2016
Rhizorhina serolis Green, 1959
Rhizorhina soyoae Kakui, 2016
Rhizorhina tanaidaceae Gotto, 1984
Rhizorhina tuberosa (Giard & Bonnier, 1893)
SPHAERONELLA Salensky, 1868
Sphaeronella abyssi Hansen, 1897
Sphaeronella acanthozonis Hnasen, 1897
Sphaeronella aeginae Hansen, 1923
Sphaeronella affinis Hansen, 1897
Sphaeronella amphilochi Hansen, 1897
Sphaeronella anarthronis Bradford, 1975
Sphaeronella antarctica Bradford, 1975
Sphaeronella antillensis Hansen, 1897
Sphaeronella argissae Hansen, 1897
Sphaeronella atyli Hansen, 1897
Sphaeronella australis Boxshall & Harrison, 1988
Sphaeronella bonnieri Hansen, 1897
Sphaeronella bouvieri Hansen, 1897
Sphaeronella bradfordae Boxshall & Lincoln, 1983
Sphaeronella calliopii Hansen, 1897
Sphaeronella callisomae Scott T., 1904
Sphaeronella capensis Hansen, 1897
Sphaeronella caprellae Blake, 1929
Sphaeronella chinensis Hansen, 1897
Sphaeronella cluthae Scott T., 1904
Sphaeronella curtipes Hansen, 1897
Sphaeronella danica Hansen, 1897
Sphaeronella decorata Hansen, 1897
Sphaeronella devosae Stock, 1960
Sphaeronella diadema Giard & Bonnier, 1895
Sphaeronella dikrothrix (Kornicker & Bowman, 1969)
Sphaeronella dispar Hansen, 1897
Sphaeronella doloriae Bradford, 1975
Sphaeronella dulichiae Hansen, 1897
Sphaeronella ecaudata Stock, 1960
Sphaeronella frontalis Hansen, 1897
Sphaeronella giardii Hansen, 1897
Sphaeronella gitanopsidis Hansen, 1897
Sphaeronella gottoi O'Reilly, 2001
Sphaeronella grayi Bradford, 1975
Sphaeronella hebe (Bowman & Kornicker, 1968)
Sphaeronella holboelli Hansen, 1897
Sphaeronella homasteropeae Bradford, 1975
Sphaeronella insignis Hansen, 1897
Sphaeronella intermedia Hansen, 1897
Sphaeronella irregularis Hansen, 1897
Sphaeronella keppelensis O'Reilly, 2002
Sphaeronella kornickeri Bradford, 1975
Sphaeronella leptocheiri Hansen, 1897
Sphaeronella leuckarti Salensky, 1868
Sphaeronella longipes Hansen, 1897
Sphaeronella marginata Hansen, 1897
Sphaeronella melphidippae Hansen, 1923
Sphaeronella messinensis Hansen, 1897
Sphaeronella metavargulae Bradford, 1975
Sphaeronella metopae Hansen, 1897
Sphaeronella microcephala Giard & Bonnier, 1893
Sphaeronella minuta Scott T., 1904
Sphaeronella modesta Hansen, 1897
Sphaeronella monothrix (Bowman & Kornicker, 1967)
Sphaeronella munnae Hansen, 1923
Sphaeronella munnopsidis Hansen, 1897
Sphaeronella nannonisci Hansen, 1923
Sphaeronella norvegica Hansen, 1904
Sphaeronella paradoxa Hansen, 1897
Sphaeronella parasteropeae Bradford, 1975
Sphaeronella philomedesi Bradford, 1975
Sphaeronella photidis Blake, 1929
Sphaeronella pikei Green 19508 (?)
Sphaeronella pilosa Blake, 1929
Sphaeronella pleurogonii Hansen, 1923
Sphaeronella pygmaea Scott T., 1904
Sphaeronella rotundata Hansen, 1923
Sphaeronella rugosidoloriae Bradford, 1975
Sphaeronella sarsiellae Bradford, 1975
Sphaeronella scleroconchae Bradford, 1975
Sphaeronella sedentaria Bonnier, 1896 (syn. Homoeoscelis sedentaria (Bonnier, 1898)
Sphaeronella serolis Monod, 1930
Sphaeronella siphonostrae Bradford, 1975
Sphaeronella skogsbergiellae Bradford, 1975
Sphaeronella spinosa Bradford, 1980
Sphaeronella squamosa Yoo & Lim, 1996
Sphaeronella synasteropeae Bradford, 1975
Sphaeronella uyenoi Kakui & Munakata, 2022 [Japan]
Sphaeronella valida Scott T., 1905
Sphaeronella vararensis Scott T., 1905
Sphaeronella vestita Hansen, 1897
SPHAERONELLOIDES Bradford, 1975
Sphaeronelloides vargulae Bradford, 1975
SPHAERONELLOPSIS Hansen, 1904
Sphaeronellopsis dikrothrix Kornicker & Bowman, 1969 (syn. Sphaeronella dikrothrix (Kornicker & Bowman, 1969)
Sphaeronellopsis hebe Bowman & Kornicker, 1968 (syn. Sphaeronella hebe (Bowman & Kornicker, 1968)
Sphaeronellopsis littoralis Hansen, 1904
Sphaeronellopsis monothrix Bowman & Kornicker, 1967 (syn. Sphaeronella monothrix (Bowman & Kornicker, 1967)
STENOTHOCHERES Hansen, 1897
Stenothocheres bruselii Hansen, 1897
Stenothocheres egregius Hansen, 1897
Stenothocheres sarsii Hansen, 1897
VUNICOTHOE Boyko, 2009
Vunicothoe cladocera (Carton, 1970)
Heptalobus Nierstrasz & Brender ŕ Brandis, 1930 syn. Choniostoma Hansen, 1886; Lecithomyzon Bloch & Gallien, 1933 syn. Choniosphaera Connolly, 1929; Pseudonicothoe Avdeev & Avdeev, 1978 syn. Paranicothoe Carton, 1970; Salenskya Giard & Bonnier, 1893 syn. Rhizorhina Hansen, 1892; Sthenothocheres syn. Stenothocheres Hansen, 1897
KEY TO GENERA OF NICOTHOIDAE
PANDARIDAE Milne Edwards, 1840 (syn. Amaterasidae Izawa, 2008 ; Cecropidae Dana, 1852; Cecropidae Yamaguti, 1963
ACHTHEINUS Wilson C.B., 1908
Achtheinus dentatus Wilson C.B., 1911
Achtheinus oblongus Wilson C.B., 1908
Achtheinus pinguis Wilson C.B., 1912
AMATERASIA Izawa, 2008
Amaterasia amanoiwatoi Izawa, 2008
Amaterasia lewisi Tang, Benz & Nagasawa, 2012
CECROPS Leach, 1816 (syn. Oniscus Lamartiniere, 1781)
Cecrops desmaresti Risso, 1816
Cecrops fulgens (Telesius, 1819)
Cecrops latreillii Leach, 1816
Cecrops lutosus (Slabber, 1760)
DEMOLEUS Heller, 18657
Demoleus ferox (Krřyer, 1838) (syn. Dinemoura ferox (Krřyer, 1838)
Demoleus heptapus (Otto, 1821)
Demoleus latus Shiino, 1954
Demoleus paradoxus (Otto, 1828) (syn. Demoleus heptapus (Otto, 1821)
DINEMOLEUS Cressey & Boyle, 1978
Dinemoleus indeprensus Cressey & Boyle, 1978
DINEMOURA Latreille, 1829 (syn. Dinematura Burmeister, 1832)
Dinemoura affinis Milne Edwards, 1840 (syn. Echthrogaleus coleoptratus (Guérin-Méneville, 1837)
Dinemoura alata Milne Edwards, 1836 (syn. Echthrogaleus coleoptratus (Guérin-Méneville, 1837)
Dinemoura braccata (Dana, 1852) (syn. Echthrogaleus coleoptratus (Guérin-Méneville, 1837)
Dinemoura coleoptrata (Guérin-Méneville, 1837) (syn. Echthrogaleus coleoptratus (Guérin-Méneville, 1837)
Dinemoura discrepans Cressey, 1967
Dinemoura ferox (Krřyer, 1838)
Dinemoura gracilis (Burmeister, 1833) (syn. Nogaus gracilis (Burmeister, 1833)
Dinemoura indistincta (Krřyer, 1863) (syn. Echthrogaleus indistinctus (Krřyer, 1863)
Dinemoura latifolia (Steenstrup & Lütken, 1861)
Dinemoura neozealanica (Thomson G.M., 1890) (syn. Echthrogaleus denticulatus Smith S.I., 1873
Dinemoura producta (Müller O.F., 1785)
Dinemoura serrata Krřyer, 1863 (syn. Philorthragoriscus serratus (Krřyer, 1863)
Dinemoura sexsetacea (Burmeister, 1833) (syn. Demoleus heptapus (Otto, 1821)
Dinemoura thynni (Krřyer, 1863) (syn. Euryphorus brachypterus (Gerstaecker, 1853)
ECHTHROGALEUS Steenstrup & Lütken, 1861
Echthrogaleus asiaticus Ho, Liu & Lin, 2012
Echthrogaleus coleoptratus (Guérin-Méneville, 1837)
Echthrogaleus denticulatus Smith S.I., 1873
Echthrogaleus disciarai Benz & Deets, 1987
Echthrogaleus mitsukurinae Izawa, 2012
Echthrogaleus pectinatus Kirtisinghe, 1964 (syn. Prosaetes rhinodontis (Wright E.P., 1876)
Echthrogaleus pellucidus Shiino, 1963
Echthrogaleus spinulus Morales-Serna, Crow, Montes & González, 2019
Echthrogaleus torpedinis Wilson C.B., 1907
ENTEPHERUS Bere, 1936
Entepherus laminipes Bere, 1936
GANGLIOPUS Gerstaecker, 1854
Gangliopus japonicus Shiino, 1960 (syn. Pannosus japonicus (Shiino, 1960)
Gangliopus pyriformis Gerstaecker, 1854
KABATAIA Kazachenko, Korotaeva & Kurochkin, 1972
Kabataia ostorhynchi Kazachenko, Korotaeva & Kurochkin, 1972
LEPIMACRUS Hesse, 1883
Lepimacrus jourdaini Hesse, 1883
LUETKENIA Claus, 1864 (syn. Cecropsina Heller, 1865)
Luetkenia asterodermi Claus, 1864
Luetkenia elongata Shiino, 1963
NESIPPUS Heller, 1865 (syn. Specilligus Dana, 1849)
Nesippus borealis Steenstrup & Lütken, 1861 (syn. Nogagus borealis Steenstrup & Lutken, 1861
Nesippus costatus Wilson C.B., 1924 (syn. Amaterasia lewisi Tang, Benz & Nagasawa, 2012
Nesippus crypturus Heller, 1865
Nesippus gonosaccus Heegaard, 1943
Nesippus nana Cressey, 1970
Nesippus orientalis Heller, 1865
Nesippus tigris Cressey, 1967
Nesippus vespa Cressey, 1964
ORTHAGORISCICOLA Poche, 1902 (syn. Laemargus Krřyer, 1837)
Orthagoriscicola muricata (Krřyer, 1837)
PAGINA Cressey, 1963
Pagina tunica Cressey, 1964
PANDARUS Leach, 1816
Pandarus alatus Johnston, 1836 (syn. Echthrogaleus coleoptratus (Guérin-Méneville, 1837)
Pandarus ambiguus (Scott T., 1907)
Pandarus bicolor Leach, 1816
Pandarus brevicaudis Dana, 1852
Pandarus carcharhini Ho, 1963
Pandarus cranchii Leach, 1819
Pandarus floridanus Cressey, 1967
Pandarus katoi Cressey, 1967
Pandarus lamnae Johnston, 1835 (syn. Dinemoura producta (Müller O.F., 1785)
Pandarus longus Gnanamuthu, 1951 (syn. Pseudopandarus longus (Gnanamuthu, 1951)
Pandarus niger Kirtisinghe, 1950
Pandarus rhincodonicus Norman, Newbound & Knott, 2000
Pandarus rouxii Risso, 1826
Pandarus satyrus Dana, 1849
Pandarus sinuatus Say, 1818
Pandarus smithii Rathbun, 1880
Pandarus unicolor Hesse, 1883 (syn. Nogaus unicolor (Hesse, 1883)
Pandarus zygaenae Brady, 1883
PANNOSUS Cressey, 1967
Pannosus japonicus (Shiino, 1960)
PARANESIPPUS Shiino, 1955
Paranesippus incisus Shiino, 1955
PERISSOPUS Steenstrup & Lütken, 1861( syn. Chlamys Beneden, 1892; Lepidopus Dana, 1849; Pholidopus Wilson C.B., 1907; Scarcina Rafinesque, 1810
Perissopus chinensis (Thomsen, 1949) accepted as Achtheinus pinguis Wilson C.B., 1912
Perissopus communis Rathbun (?)
Perissopus dentatus Steenstrup & Lütken, 1861
Perissopus galeorhini (Yamaguti, 1936) (syn. Achtheinus pinguis Wilson C.B., 1912
Perissopus impenderus (Shen & Wang, 1959) (syn. Achtheinus pinguis Wilson C.B., 1912
Perissopus intermedius (Kurtz, 1924) (syn. Achtheinus pinguis Wilson C.B., 1912
Perissopus japonicus (Wilson C.B., 1922) (syn. Achtheinus pinguis Wilson C.B., 1912
Perissopus oblongatus (Wilson C.B., 1908) (syn. Achtheinus oblongus Wilson C.B., 1908
Perissopus parvidens (Wilson C.B., 1923) (syn. Achtheinus pinguis Wilson C.B., 1912
Perissopus pinguis (Wilson C.B., 1912) (syn. Achtheinus pinguis Wilson C.B., 1912
Perissopus platensis (Thomsen, 1949) (syn. Achtheinus pinguis Wilson C.B., 1912
PHILORTHRAGORISCUS Horst, 1897
Philorthragoriscus serratus (Krřyer, 1863)
PHYLLOTHYREUS Norman, 1903 (syn. Laminifera Poche, 1902; Parapandarus Wilson C.B., 1924; Phyllophora Milne Edwards H., 1840.
Phyllothyreus cornutus (Milne Edwards, 1840)
PROSAETES Wilson C.B., 1907 (syn. Stasiotes Wright E.P., 1876)
Prosaetes rhinodontis (Wright E.P., 1876)
PSEUDOPANDARUS Kirtisinghe, 1950
Pseudopandarus australis Cressey & Simpfendorfer, 1988
Pseudopandarus cairae Bernot & Boxshall, 2017
Pseudopandarus gracilis Kirtisinghe, 1950
Pseudopandarus longus (Gnanamuthu, 1951)
Pseudopandarus pelagicus Rangnekar, 1977
Pseudopandarus scyllii Yamaguti & Yamasu, 1959
Cecropsina Heller, 1865 syn. Luetkenia Claus, 1864; Chlamys Beneden, 1892 syn. Perissopus Steenstrup & Lütken, 1861; Dinematura Burmeister, 1832 syn. Dinemoura Latreille, 1829; Laemargus Krřyer, 1837 syn. Orthagoriscicola Poche, 1902 ; Laminifera Poche, 1902 syn. Phyllothyreus Norman, 1903; Lepidopus Dana, 1849 syn.Perissopus Steenstrup & Lütken, 1861; Nessipus Heller, 1865 syn. Nesippus Heller, 1865; Nogagus Leach, 1816 syn. Nogaus Leach, 1819; Oniscus Lamartiniere, 1781 syn.Cecrops Leach, 1816; Orthagorisicola Poche, 1902 yn. Orthagoriscicola Poche, 1902; Parapandarus Wilson C.B., 1924 yn. Phyllothyreus Norman, 1903; Philorthagoriscus Horst, 1897 yn.Philorthragoriscus Horst, 1897; Pholidopus Wilson C.B., 1907 syn. Perissopus Steenstrup & Lütken, 1861; Phyllophora Milne Edwards H., 1840 syn. Phyllothyreus Norman, 1903; Specilligus Dana, 1849 syn. Nesippus Heller, 1865; Stasiotes Wright E.P., 1876 syn. Prosaetes Wilson C.B., 1907. Key to the valid species of Pseudopandarus based
on adult females
PENNELLIDAE Burmeister, 1835
(syn. Lernaeocerinae Wilson C.B., 1917; Pennellinae Wilson C.B., 1917; Pennillidae Burmeister, 1816; Perodermatidae Yamaguti, 1963; Pontopsyllidae Scott T., 1894.
"The family Pennellidae comprises ecto and mesoparasitic copepods on marine fishes.
Although a preliminary scheme of phylogenetic
relationships of pennellids based on morphological
characters exists, it is difficult to objectively define
character states because of their highly modified
bodies and reduced appendages.
In the family Pennellidae, 25 genera and ca. 180
species have been described from fishes and occasionally from marine mammals in the world's oceans"
(From Yumura et al., 2021)
PSEUDOSARCOTRETES Yumura et al. 2024
Pseudosarcotretes omorii Yumura et al. 2024 [Japan]
ALLOTRIFUR Yamaguti, 1963
Allotrifur chlorophthalmi (Yamaguti, 1939)
CARDIODECTES Wilson C.B., 1917 "Currently, Cardiodectes comprises 16 species, which are separated into two morphological groups, 'medusaeus' and 'rubosus' (Izawa, 1970; Bellwood, 1981). The former contains the following 5 species, Cardiodectes anchorellaeBrian and Gray, 1928, C. bellottii (Richiardi, 1882), C. cristatus Shiino, 1958, C. frondosus Schuurmans-Stekhoven, 1937, C. longicervicus Shiino, 1958, and the group is recognized by possessing a urosome. The second group comprises the remaining 11 species; C. asper Uyeno and Nagasawa, 2010, C. bertrandi Uyeno and Nagasawa, 2010, C. bellwoodi Uyeno, 2013, C. boxshalli Bellwood, 1981, C. hardenbergi Markevich, 1936, C. krishnai Sebastian, 1968, C. rotundicaudatus Izawa, 1970, C. rubosus Leigh-Sharpe, 1934, C. shini Uyeno, 2013, C. spiralis Bellwood, 1981, and C. roatanenis Suárez-Morales, Vásquez-Yeomans and Vidotto, 2022 (Izawa, 1970; Bellwood, 1981; Suarez- Morales et al., 2022" (from Aneesh et al. 2023
Cardiodectes anchorellae Brian & Gray, 1928
Cardiodectes asper Uyeno & Nagasawa, 2010
Cardiodectes bellottii (Richiardi, 1882)
Cardiodectes bellwoodi Uyeno, 2013
Cardiodectes bertrandi Uyeno & Nagasawa, 2010
Cardiodectes boxshalli Bellwood, 1981
Cardiodectes cristatus Shiino, 1958
Cardiodectes frondosus Schuurmans Stekhoven J.H. Jr, 1937
Cardiodectes hardenbergi Markevich, 1936
Cardiodectes krishnai Sebastian, 1968
Cardiodectes longicervicus Shiino, 1958
Cardiodectes medusaeus (Wilson C.B., 1908) (syn. Cardiodectes bellottii (Richiardi, 1882)
Cardiodectes roatanensis Suárez-Morales, Vásquez-Yeomans & Vidotto, 2022
Cardiodectes rotundicaudatus Izawa, 1970
Cardiodectes rubosus Leigh-Sharpe, 1934
Cardiodectes shini Uyeno, 2013
Cardiodectes spiralis Bellwood, 1981
Cardiodectes vampire Aneesh et al.m 2023 [Indian Ocean]
KEY TO SPECIES OF CARDIODECTES (FEMALES)
CREOPELATES Shiino, 1958
Creopelates floridus Shiino, 1958
Creopelates hosinoi Uyeno, 2015
Creopelates lubangenesis Uyeno, 2015
Creopelates nohmijimensis Uyeno & Nagasawa, 2010
Creopelates shirakawai Uyeno, 2015
EXOPENNA Boxshall, 1986
Exopenna crimmeni Boxshall, 1986
HAEMOBAPHES Steenstrup & Lütken, 1861 (SYN.Anops Oken, 1815; Collipravus Wilson C.B., 1917,Haemobaphoides Scott T. & Scott A., 1913)
Haemobaphes ambiguus Scott T., 1900
Haemobaphes cresseyi Kazachenko, 1995
Haemobaphes cyclopterina (Müller O.F., 1776)
Haemobaphes diceraus Wilson C.B., 1917
Haemobaphes disphaerocephalus Grabda, 1976
Haemobaphes enodis Wilson C.B., 1917
Haemobaphes intermedius Kabata, 1967
Haemobaphes pannosus Kabata, 1979
Haemobaphes parvus (Wilson C.B., 1917)
Haemobaphes puntaarenensis Muńoz & Castro, 2022 [South Chile]
IMPEXUS Kabata, 1972
Impexus hamondi Kabata, 1972
LERNAEENICUS Lesueur, 1824 (syn. Foroculum Thompson W., 1844; Lernaeonema Milne Edwards H., 1840; Lerneoceropsis Fowler, 1912)
Lernaeenicus affixus Wilson C.B., 1917
Lernaeenicus alatus Rangnekar, 1961
Lernaeenicus anchoviellae Sebastian & George, 1964
Lernaeenicus ater Shiino, 1958
Lernaeenicus bataviensis Sebastian, 1967
Lernaeenicus centropristi (Pearse, 1947)
Lernaeenicus cerberus Leigh-Sharpe, 1927
Lernaeenicus encrasicholi (Turton, 1807)
Lernaeenicus eristaliformis Brian, 1908 (syn. Sarcotretes eristaliformis (Brian, 1908)
Lernaeenicus gempyli Horst, 1878 (syn. Sarcotretes scopeli Jungersen, 1911
Lernaeenicus gnathonicus Leigh-Sharpe, 1934
Lernaeenicus gnavus Leigh-Sharpe, 1934 (syn. Protosarcotretes gnavus (Leigh-Sharpe, 1934)
Lernaeenicus gonostomae Kensley & Grindley, 1973
Lernaeenicus gracilis (Heller, 1865)
Lernaeenicus hemirhamphi Kirtisinghe, 1932
Lernaeenicus kabatai Oldewage, 1989
Lernaeenicus inflexus Steenstrup & Lütken, 1861 (syn. Sarcotretes scopeli Jungersen, 1911)
Lernaeenicus longiventris Wilson C.B., 1917
Lernaeenicus medusaeus Wilson C.B., 1908 (syn. Cardiodectes medusaeus (Wilson C.B., 1908)
Lernaeenicus megalaspis Aneesh, Helna, Kumar & Venmathi Maran, 2021
Lernaeenicus mulli (Beneden, 1870)
Lernaeenicus multilobatus Lewis, 1959 (syn. Protosarcotretes multilobatus (Lewis, 1959)
Lernaeenicus musteli (Beneden, 1851) (syn. ripaphylus musteli (Beneden, 1851)
Lernaeenicus neglectus Richiardi, 1877
Lernaeenicus nemipteri Gnanamuthu, 1953
Lernaeenicus nodicornis Steenstrup & Lütken, 1861 (syn. Sarcotretes scopeli Jungersen, 1911
Lernaeenicus ocularis (Cuvier, 1830)
Lernaeenicus polyceraus Wilson C.B., 1917
Lernaeenicus polynemi (Bassett-Smith, 1898)
Lernaeenicus procerus (Leidy, 1889)
Lernaeenicus quadrilobatus Yamaguti & Utinomi, 1953
Lernaeenicus radiatus Lesueur, 1824
Lernaeenicus ramosus Kirtisinghe, 1956
Lernaeenicus sayori Yamaguti, 1939
Lernaeenicus seeri Kirtisinghe, 1934
Lernaeenicus sprattae (Sowerby, 1806)
Lernaeenicus stromatei Gnanamuthu, 1953
Lernaeenicus triangularis Heegaard, 1966
Lernaeenicus vorax Richiardi, 1877
LERNAEOCERA Blainville, 1822 (syn, Saucissona Leigh-Sharpe, 1935 )
Lernaeocera abyssicola (Brady, 1883)
Lernaeocera barnimiana Hartmann, 1865 (syn. Lernaea barnimii (Hartmann, 1870)
Lernaeocera branchialis (Linnaeus, 1767)
Lernaeocera caparti Machado-Cruz, 1959
Lernaeocera centropristi Pearse, 1947 (syn. Lernaeenicus centropristi (Pearse, 1947)
Lernaeocera cyclopterina Blainville, 1822 (syn. Haemobaphes cyclopterina (Müller O.F., 1776)
Lernaeocera esocina (Hermann, 1783)
Lernaeocera galei Krřyer, 1837
Lernaeocera godfroyi (Quidor, 1913) (syn. Trifur godfroyi (Quidor, 1913)
Lernaeocera lotellae (Thomson G.M., 1890) (syn. Trifur tortuosus Wilson C.B., 1917
Lernaeocera lusci (Bassett-Smith, 1896)
Lernaeocera minuta (Scott T., 1900)
Lernaeocera rigida (Krřyer, 1863) (syn. Trifur tortuosus Wilson C.B., 1917
Lernaeocera sauciatonis (Leigh-Sharpe, 1935)
Lernaeocera surriraiis Blainville, 1822 (syn. Lernaeenicus sprattae (Sowerby, 1806)
LERNAEOLOPHUS Heller, 1865
Lernaeolophus aceratus Ho & Honma, 1983
Lernaeolophus recurvus Wilson (?)
Lernaeolophus striatus Wilson C.B., 1913
Lernaeolophus sultanus (Milne Edwards, 1840)
METAPENICULUS Castro-Romero & Baeza-Kuroki, 1985
Metapeniculus antofagastiensis Castro-Romero & Baeza-Kuroki, 1985
Metapeniculus haemuloni (Alexander, 1983)
NAGASAWANUS Uyeno, 2015
Nagasawanus akinohama Uyeno, 2015
Nagasawanus snufkini Uyeno, 2015
OPHIOLERNAEA Shiino, 1958
Ophiolernaea formosana Ho, 1966
Ophiolernaea longiceps Shiino, 1958
PARINIA Kazachenko & Avdeev, 1977
Parinia myctophi Kazachenko & Avdeev, 1977
PENICULISA Wilson C.B., 1917
Peniculisa bellwoodi Boxshall, 1989
Peniculisa bicaudata Shiino, 1956
Peniculisa crassa Uyeno & Nagasawa, 2010
Peniculisa elongata Uyeno & Nagasawa, 2010
Peniculisa furcata (Krřyer, 1863)
Peniculisa ohirugi Uyeno & Nagasawa, 2010
Peniculisa shiinoi Izawa, 1965
Peniculisa uchinah Uyeno & Nagasawa, 2010
Peniculisa wilsoni Radhakrishnan, 1977
PENICULUS Nordmann, 1832
Peniculus asinus Kabata & Wilkes, 1977 (syn. Pseudopeniculus asinus (Kabata & Wilkes, 1977)
Peniculus calamus Nordmann, 1864
Peniculus clavatus Kröyer, 1863
Peniculus communis Leigh-Sharpe, 1934
Peniculus elegans Leigh-Sharpe, 1934
Peniculus elongatus Boxshall, 1986
Peniculus furcatus Krřyer, 1863 (syn. Peniculisa furcata (Krřyer, 1863)
Peniculus haemuloni Alexander, 1983 (syn. Metapeniculus haemuloni (Alexander, 1983)
Peniculus hokutoae Ohtsuka, Nishikawa & Boxshall, 2018
Peniculus minuticaudae Shiino, 1956
Peniculus ostraciontis Yamaguti, 1939
Peniculus quadratus Moon & Choi, 2014
Peniculus sciaenae Gnanamuthu, 1951 (syn. Propeniculus
Peniculus scomberi Gnanamuthu, 1951 (syn. Propeniculus scomberi (Gnanamuthu, 1951)
Peniculus stromatei Gnanamuthu, 1951 (syn. Propeniculus stromatei (Gnanamuthu, 1951)
Peniculus truncatus Shiino, 1956
Peniculus theraponi Gnanamuthu, 1951 (syn. Propeniculus theraponi (Gnanamuthu, 1951)
Peniculus trichiuri Gnanamuthu, 1951 (syn. Propeniculus trichiuri (Gnanamuthu, 1951)
PENNELLA Oken, 1815
(syn.Baculus Lubbock, 1860; Hessella Brady, 1883; Lernaeopenna Blainville, 1822; Lerneopenna Blainville, 1822;
Penella (Oken) Quidor, 1913;
Penicillus Kumar & Hameed, 1993.
Pennella balaenoptera Koren & Danielssen, 1877
Pennella benzi Hogans, 2017
Pennella diodontis Oken, 1815
Pennella exocoeti (Holten, 1802)
Pennella filosa (Linnaeus, 1758)
Pennella hawaiiensis Kazachenko & Kurochkin, 1974
Pennella instructa Wilson C.B., 1917
Pennella makaira Hogans, 1988
Pennella remorae Murray, 1856
Pennella sagitta (Linnaeus, 1758)
Pennella sultana Milne-Edwards, 1840 (syn. Lernaeolophus sultanus (Milne Edwards, 1840)
PERODERMA Heller, 1865 (syn. Taphrobia Cornalia, 1875)
Peroderma bellotti Richiardi, 1882 (syn. Cardiodectes bellottii (Richiardi, 1882)
Peroderma cylindricum Heller, 1865
Peroderma pacifica Izawa, 1977
Peroderma petersi Richiardi, 1881
Peroderma sardinellae Ranjit & Bensam, 1998
Peroderma tasselum Bennet & Chellam, 1975
PHRIXOCEPHALUS Wilson C.B., 1908 (syn. Phrixocephaloides Yamaguti, 1963)
Phrixocephalus alatus Shiino, 1956
Phrixocephalus arboreus Shiino, 1956
Phrixocephalus carcellesi Brian, 1944
Phrixocephalus cepolae Yamaguti, 1939
Phrixocephalus cincinnatus Wilson C.B., 1908
Phrixocephalus crassus Shiino, 1956
Phrixocephalus curvatus Shiino, 1956
Phrixocephalus diversus Wilson C.B., 1917
Phrixocephalus longicollum Shiino, 1956
Phrixocephalus reniformis Shiino, 1956
Phrixocephalus triangulus Wilson C.B., 1917
Phrixocephalus umbellatus Shiino, 1956
Phrixocephalus vipereus Shiino, 1956
PROPENICULUS Castro Romero, 2014
Propeniculus sciaenae (Gnanamuthu, 1951) (syn. Propeniculus trichiuri Gnanamuthu, 1951)
Propeniculus scomberi (Gnanamuthu, 1951) (syn. Propeniculus trichiuri Gnanamuthu, 1951)
Propeniculus stromatei (Gnanamuthu, 1951) (syn. Propeniculus trichiuri Gnanamuthu, 1951)
Propeniculus theraponi (Gnanamuthu, 1951) (syn. Propeniculus trichiuri Gnanamuthu, 1951)
Propeniculus trichiuri (Gnanamuthu, 1951)
PROTOSARCOTRETES Ohtsuka, Lindsay & Izawa, 2018
Protosarcotretes gnavus (Leigh-Sharpe, 1934)
Protosarcotretes multilobatus (Lewis, 1959)
Protosarcotretes nishikawai Ohtsuka, Lindsay & Izawa, 2018
PSEUDOPENICULUS Castro Romero, 2014
Pseudopeniculus asinus (Kabata & Wilkes, 1977)
SARCOTRETES Jungersen, 1911
Sarcotretes eristaliformis (Brian, 1908)
Sarcotretes gonostomae n. comb. Sebone & Dippenaar 2024
Sarcotretes longirostris Ho, Nagasawa & Kim I.H., 2007
Sarcotretes scopeli Jungersen, 1911
Sarcotretes umitakae Uyeno, Wakabayashi & Nagasawa, 2012
SERPENTISACCUS Blasiola, 1979
Serpentisaccus magnificus Blasiola, 1979
TRIFUR Wilson C.B., 1917
"The genus Trifur Wilson, 1917 comprises relatively large and
conspicuous parasitic copepods infecting the body surface and
gills of several marine fishes. It was erected to accommodate
T. tortuosus Wilson, 1917, a parasite of the body surface of
Salilota australis (Ghnther, 1878) from the Smith Channel,
Otter Bay Southern Chile (Wilson 1917). Later on, other
species were described from the Southern Hemisphere:
T. merluccii Talice, 1936, parasitizing “Merluccius gayi
(Guichenot, 1848)”, actually Merluccius hubbsi Marini, 1933
from Uruguay (Talice 1936); T. puntaniger Thomé, 1963 from
an unknown host species caught in the coasts of Uruguay
(Thomé 1963); T. chlorophthalmi Yamaguti, 1939 from Chlorophthalmus
albatrossis Jordan et Starks, 1904 from Japan
(Yamaguti 1963) and T. physiculi Heegaard, 1962 from the
skin of “Physiculus barbatus (Ghnther, 1863)” = Pseudophysis
barbata Günther, 1863 and “Physiculus sp.” from Australia
and Tasmania (Heegaard 1962)" (from: Etchegoin et al., 2009)
Trifur chlorophthalmi Yamaguti, 1939 (syn. Allotrifur chlorophthalmi (Yamaguti, 1939)
Trifur godfroyi (Quidor, 1913)
Trifur merluccii Talice, 1936 (?)
Trifur tortuosus Wilson C.B., 1917
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