GISELINIDAE Martinez Arbizu, 2000

GISELINA Martinez Arbizu, 2000

Type-species: Giselina cristata Martinez Arbizu, 2000

  • Giselina cristata Martinez Arbizu, 2000 [Atlantic Ocean]
  • Giselina gerdesi Martinez Arbizu, 2000 [Atlantic Ocean]

    SENSOGISELINA Martinez Arbizu, 2000

    Type-species: Sensogiselina potteki Martinez Arbizu, 2000

  • Sensogiselina potteki Martinez Arbizu, 2000 [Atlantic Ocean]
  • Sensogiselina reducta Martinez Arbizu, 2000 [Atlantic Ocean]

    MANTRIDAE Leigh-Sharpe, 1934

    CHAMICOLA Ohtsuka, Boxshall & Torigoe, 2000

  • Chamicola nagasawai Ohtsuka, Boxshall & Torigoe, 2000

    MANTRA Leigh-Sharpe, 1934

  • Mantra speciosa Leigh-Sharpe, 1934

    NEOARCHINOTODELPHYS Ummerkutty, 1961

  • Nearchinotodelphys indicus Ummerkutty, 1961




    PONTOSTRATOTIDAE Scott A., 1909

    According to Huys 2009, the family-group name Pontostratiotidae A. Scott, 1909 (type: Pontostratiotes Brady, 1883) is a senior subjective synonym of Cerviniopseinae Brotskaya, 1963 (type: Cerviniopsis Sars, 1903) and the former is consequently reinstated at the subfamilial level.

  • Pontostratiotes abyssicola Brady, 1883




    POECILOSTOMATOIDA

    Poecilostomatoida are a suborder of copepods. Although it was previously considered a separate order, recent research showed it to be nested within the Cyclopoida (Khodami et al., 2017), and most families formerly included in Poecilostomatoida are now included in the suborder Ergasilida (Khodami et al., 2019)


    CLAUSIDIIDAE Embleton, 1901

    "Copepods of the family Clausidiidae Embleton, 1901 represent an early offshoot of the poecilostome lineage within the order Cyclopoida Burmeister, 1834 (see Khodami et al., 2017). Most clausidiids live in loose association with marine invertebrate hosts (Huys & Boxshall, 1991) and species of Clausidium Kossmann, 1874 are recorded exclusively living in association with ghost shrimps (Boxshall & Halsey, 2004)"

    Key to genera of Clausidiidae


    HYPHALION Humes, 1987

  • Hyphalion captans Humes, 1987
  • Hyphalion sagamiense Toda, Miura & Nemoto, 1992
  • Hyphalion tertium Defaye & Toda, 1994


    LEPTINOGASTER Pelseneer, 1929 (syn. Myocheres Wilson C.B., 1950; Strongylopleura Pelseneer, 1929)

  • Leptinogaster californiensis Gooding, 1963
  • Leptinogaster dentata (Humes & Cressey, 1958)
  • Leptinogaster digita Kim I.H. & Ho, 1991
  • Leptinogaster histrio (Pelseneer, 1929)
  • Leptinogaster inflata (Allen, 1956)
  • Leptinogaster major (Williams, 1907)
  • Leptinogaster minuta Kim I.H., 2009
  • Leptinogaster pholadis Pelseneer, 1929
  • Leptinogaster pruvoti (Monod & Dollfus, 1932)
  • Leptinogaster scobina (Humes & Cressey, 1958)
  • Leptinogaster stocki Ranade, 1974


    IDOMENE Philippi, 1843 (taxon inquirendum)

    HERSILIA Philippi, 1839 accepted as Clausidium Kossmann, 1874 (preoccupied by Hersilia Savigny, 1826 (Arachnida)

    MYOCHERES Wilson C.B., 1950 accepted as Leptinogaster Pelseneer, 1929 (synonym)

    PLATYCHEROS Scott T., 1892 accepted as Hemicyclops Claus, 1893

    PONTOCYCLOPS Serban, 1956 accepted as Hemicyclops Boeck, 1872 (synonym)

    SAPHIRELLA Scott T., 1894 syn.Lanowia Oliveira, 1945; Sapphirella Bjornberg, 1963 (Clausidiidae Embleton, 1901 1901)

    STRONGYLOPLEURA Pelseneer, 1929 accepted as Leptinogaster Pelseneer, 1929 (synonym)

    TOCOCHERES Pelseneer, 1929 accepted as Hemicyclops


    HEMICYCLOPS Boeck, 1872 (syn. Pontocyclops Serban, 1956)

    "The cyclopoid copepod genus Hemicyclops accommodates about 40 species from shallow waters of the world (Kim 2000; Boxshall and Halsey 2004; Mulyadi 2005), Recently ,Kim (2007)ransferre dfbur species to the new genus Goodingius .The development and life cycle of Hemicyclops), as well investigated Ito & Nishida 1991 ,1995 ,1997, 2007, 2008; Kim and Ho 1992 ;Itoh 2001) .Six naupliar and the first copepodid stages are free-swimming and serve for dispersa la,nd then the first copepodid stage settles on the bottom; the second and subsequent copepodid stages and adults are loosely associated with benthic animals such as polychaetes, echiurans, crabs, and mud shrimps. The firs ctopepodid stage was previousl yreferred to as Sqphirelt abased on its unique morphology and predominance in plankton (Itoh and Nishida 1991; Itoh 2001)" (from: Ohtsuka et al., 2010)

    "Several species of Hemicyclops Boeck, 1873, a genus of copepods widely distributed in temperate and tropical regions of the world, are associated with various invertebrates (a sponge, a gastropod, a bivalve, polychaetes and crustaceans). Other species have been found in burrows of polychaetes or crustaceans and are suspected to have associations with such hosts. For still other species, regarded as free-living, no associations are known. In this paper a new species of Hemicyclops is reported for the first time from a hard coral" (from: Humes, 1984)

  • Hemicyclops aberdonensis O' Reilly, 2000 [Indonesia]
  • Hemicyclops acanthophorus Humes, 1995 [Madagascar]
  • Hemicyclops acanthosquillae Humes, 1965
  • Hemicyclops amplicaudatus Humes, 1965
  • Hemicyclops adhaederens (Williams) (?)
  • Hemicyclops apiculus Humes, 1995 [Madagascar]
  • Hemicyclops australis Nicholls, 1944
  • Hemicyclops axiophilus Humes, 1965
  • Hemicyclops biflagellatus Humes, 1965
  • Hemicyclops columnaris Humes, 1984
  • Hemicyclops cornutus Kim I.H. & Hong, 2014
  • Hemicyclops caissarum Kikara & Da Rocha, 1993 [Brazil] (syn. Giardella caissarum (Kihara & Rocha, 1993)
  • Hemicyclops carinifer Humes, 1965 (syn. Giardella carinifer (Humes, 1965)
  • Hemicyclops ctenidis Ho & Kim I.H., 1990
  • Hemicyclops cylindraceus (Pelseneer, 1929)
  • Hemicyclops dilatatus Shen & Bai, 1956
  • Hemicyclops diremptus Humes, 1965
  • Hemicyclops dubia (Thompson I.C. & Scott A., 1903)
  • Hemicyclops geminatus Stock, 1992 [Curacao] (?)
  • Hemicyclops giardelloides Gooding, 1963
  • Hemicyclops gomsoensis O' Reilly, 2000 [Indonesia]
  • Hemicyclops humesi Kim I.H., 2009
  • Hemicyclops indicus Sewell, 1949
  • Hemicyclops intermedius Ummerkutty, 1962
  • Hemicyclops membranatus Moon & Kim, 2010 [Korea]
  • Hemicyclops minutus Mulyadi.. 2005 [Indonesia]
  • Hemicyclops nasutus Moon & Kim, 2010 [Korea]
  • Hemicyclops nichollsi Karanovic, 2008)
  • Hemicyclops japonicus Itoh & Nishida,1993 [Japan]
  • Hemicyclops javanensis Mulyadi.. 2005 [Indonesia]
  • Hemicyclops kombensis Humes, 1965
  • Hemicyclops leggii (Thompson I.C. & Scott A., 1903)
  • Hemicyclops livingstoni (Scott T., 1894)
  • Hemicyclops magnus Kim I.H., 2009
  • Hemicyclops membranatus Moon & Kim I.H., 2010
  • Hemicyclops minutus Mulyadi, 2005
  • Hemicyclops mortoni Boxshall & Humes, 1987
  • Hemicyclops nasutus Moon & Kim I.H., 2010
  • Hemicyclops nichollsi Karanovic, 2008
  • Hemicyclops parapiculus Kim I.H. & Hong, 2014
  • Hemicyclops parilis Moon & Kim, 2010 [Korea]
  • Hemicyclops perinsignis Humes, 1973
  • Hemicyclops planifusus Boeck, 1972
  • Hemicyclops puffini (Thompson I.C., 1888)
  • Hemicyclops purpureus Boeck, 1872
  • Hemicyclops rapax Lee et. al., 2022 [Korea]
  • Hemicyclops saxatilis Ho & Kim I.H., 1991
  • Hemicyclops sebastiani Kikara & Da Rocha, 1993 [Brazil]
  • Hemicyclops spinulosus Ito &Ishida, 1988 [Japan]
  • Hemicyclops subadhaerensis Gooding, 1960 (syn. Goodingius subadhaerens (Gooding, 1960)
  • Hemicyclops tamilensis (Thompson I.C. & Scott A., 1903)
  • Hemicyclops tanakai Itoh & Ishida, 2002 [Japan]
  • Hemicyclops thalassius Vervoort & Ramirez, 1996 [Argentina]
  • Hemicyclops thompsoni (Canu, 1888)
  • Hemicyclops thysanotus Wilson C.B., 1935
  • Hemicyclops tripartitus Kim I.H., 2009
  • Hemicyclops ventriplanus Kim I.H., 2000
  • Hemicyclops vicinalis Humes, 1995 [Madagascar]
  • Hemicyclops visendus Humes, Cressey & Gooding, 1958
  • Hemicyclops xiamenensis Ohtsuka et al., 2010 [China]


    CONCHYLIURUS Bocquet & Stock, 1957

  • Conchyliurus bombasticus Reddiah, 1961
  • Conchyliurus cardii Gooding, 1957
  • Conchyliurus dispar Moon & Kim, 2014 [Yellow Sea]
  • Conchyliurus fragilis Reddiah, 1961 (syn. Conchyliurus bombasticus Reddiah, 1961
  • Conchyliurus inchonensis Kim I.H., 1997
  • Conchyliurus lobatus Humes & Cressey, 1958
  • Conchyliurus mactrae Avdeev, 1977
  • Conchyliurus maximus Reddiah, 1961
  • Conchyliurus quintus Kim, 1994 [Korea]
  • Conchyliurus solenis Bocquet & Stock, 1957
  • Conchyliurus torosus Humes & Cressey, 1958


    HEMADOMA Kim, 2004

  • Hemadoma clavicrura Kim, 2004 [Korea]

    PSEUDANTHESSIUS Claus, 1889

  • Pseudanthessius excertus Kim, 2014 [Korea]
  • Pseudanthessius spinosus Kim, 2014 [Korea]

    SEWELLOYA Kim, 2014

  • Sewelloya plana Kim, 2014 [Korea]

    GOODINGIUS Kim, 2007

  • Goodingius adhaereas (William, 1907)
  • Goodingius elongatus Wilson, 1937
  • Goodingius arenicolae Gooding, 1960
  • Goodingius subadhaereas Gooding, 1960

    HERSILIODES Kanu, 1988

  • Hersiliodes dubia Thompson & Scott 1901 (?)
  • Hersiliodes exiguus Kim, 1996 [Korea]
  • Hersiliodes latericia (Grube, 1869)
  • Hersiliodes leggii Thompson & Scott 1901 (?)
  • Hersiliodes tamilensi Thompson & Scott 1901 (?)

    GIARDIELLA Kanu, 1888

  • Giardiella callianassae Kanu, 1888
  • Giardiella thomsoni Kanu, 1888
  • Giardiella richoensis Kim. 2007
  • Giardiella carinifer Humes, 1965
  • Giardiella caissarum Kikaka & Rocha, 1993


    CLAUSIDIUM Kossmann, 1875 (syn. Hersilia Philippi, 1839)

    Clausidium is a genus of copepods that is found in subtropical to temperate coastal areas. All species of the genus occur on the bodies of mud shrimp of the families Callianassidae and Upogebiidae.

  • Clausidium apodiformis (Philippi, 1839)
  • Clausidium californiense Wilson C.B., 1935 (syn.Clausidium vancouverensis (Haddon, 1912)
  • Clausidium caudatum (Say, 1818)
  • Clausidium chelatum Pillai, 1959
  • Clausidium dissimile Wilson C.B., 1921
  • Clausidium iranensis Sephavand, Kihara & Boxshall, 2019
  • Clausidium makranensis Sepahvand et al., 2017 [Iran; marine]
  • Clausidium maximus Hwang et al., 2016 [Korea; associate of a burrowing decapod]
  • Clausidium persiaensis Sepahvand and Kihara [Persian Gulf, Indian Ocean]
  • Clausidium rodriguezi Kikara & Da Rocha, 2013 [Brazil]
  • Clausidium saldanhae Kensley, 1974 [Africa]
  • Clausidium sarii Sepahvand et al., 2017 [Iran; marine]
  • Clausidium searsi Wilson C.B., 1937
  • Clausidium senegalense Humes, 1957
  • Clausidium tenax Humes, 1949
  • Clausidium testudo Kossmann, 1874 (syn. Clausidium apodiformis (Philippi, 1839)
  • Clausidium travancorense Pillai, 1959
  • Clausidium vancouverensis (Haddon, 1912)

    KEY TO CLAUSIDUM SPECIES


    HIPPOMOLGUS G. O. Sars, 1917

  • Hippomolgus cognatus Humes & Ho, 1967
  • Hippomolgus furcifer Sars G.O., 1917
  • Hippomolgus latipes Humes & Ho, 1967
  • Hippomolgus limiticus Hwang et al., 2016 [China; muddy bottom sediments]


    CEMIHYCLOPS Karanovic, 2008

  • Cemihyclops ceduensis Karanovic, 2008
  • Cemihyclops tenuis Kim I.H., 2009

    FOLIOMOLGUS Kim, 2001

  • Foliomolgus cucullus Kim, 2001 [Korea]


    HERILIA Philippi, 1939


    PHOLADICOLA Ho & Wardle, 1992

  • Pholadicola thomassini Lucien(?)
  • Pholadicola intestinalis Ho & Wardle, 1992

    PSEUDOERSILIA Strand, 1914


    PSEUDOPSYLLUS Scott, 1902 (taxon inquirendum, listed as genus inquirendum by Boxshall & Halsey, 2004)


    SAPHIRELLA Scott, 1804


    STOCKIA Sebastian & Pillai, 1974

  • Stockia indica Sebastian & Pillai, 1974

    HAMATICYCLOPS Kim, 2015

  • Hamaticyclops ahni Kim, 2015 [California; marine]




    CLAUSIIDAE Giesbrecht, 1895


    "Copepods of the family Clausiidae are essentially living in symbiosis with the polychaetes (O’Reilly, 1995; Kim, 2001). Although the name ‘‘Familie Clausiidae’’ appeared in the 19th century (Giesbrecht, 1895: 226), the definition of the family was not formally given until 1955 when Wilson and Illg gave an emended diagnosis of the Clausiidae. Only six species in five genera (Clausia, Mesnilia, Rhodinicola, Seridium, Teredicola) were recognized. Although 12 genera have been ‘‘put forward as tentative clausiids’’ (O’Reilly, 1995: 57), through the works of Gooding (1963), Stock (1968, 1979), and Ho (1984), by the time O’Reilly (1995) reported a new genus and new species of clausiid (Megaclausia mirabilis) from the maldanid polychaetes obtained in the Firth of Forth off the coast of Scotland, only seven genera (Clausia, Indoclausia, Mesnilia, Pontoclausia, Pseudoclausia, Rhodinicola, Stockia) were recognized as valid in the family" (from: Ho & Kim, 2003)

    LIKROCLAUSIA Ho & Kim I. H. 2003

  • Likroclausia namhaensis Ho & Kim I. H. 2003

    MAXILLICLAUSIA Kim, 2014

  • Maxilliclausia propria Kim, 2014 [Korea]

    OSHOROCLAUSIA Uyeno & Kakui, 2015

  • Oshoroclausia shibazakii Uyeno & Kakui, 2015

    VIVGOTTOIA Kim I.H., Sikorski, O'Reilly & Boxshall, 2013

  • Vivgottoia garwoodi Kim I.H., Sikorski, O'Reilly & Boxshall, 2013

    BOREOCLAUSIA Kim I.H., Sikorski, O'Reilly & Boxshall, 2013

  • Boreoclausia holmesi Kim I.H., Sikorski, O'Reilly & Boxshall, 2013
  • Boreoclausia recta Kim I.H., Sikorski, O'Reilly & Boxshall, 2013

    SHEADERIA Kim I.H., Sikorski, O'Reilly & Boxshall, 2013

  • Sheaderia bifida Kim I.H., Sikorski, O'Reilly & Boxshall, 2013

    CLAUSIA Claparčde, 1863 (syn. Doviella Rocha C.E.F., 1989)

  • Clausia antiqua Kim I.H., 2001
  • Clausia lobata Kim I.H., 2000
  • Clausia lubbockii Claparčde, 1863
  • Clausia parva Kim, 2014 [Korea]

    DONUSA Nordmann, 1864

  • Donusa clymenicola Nordmann, 1864

    FLABELLIPHILUS Bresciani & Lützen, 1962

  • Flabelliphilus inersus Bresciani & Lützen, 1962

    PONTOCLAUSIA Bacescu & Por, 1959

  • Pontoclausia cochleata Lee et al., 2022 [Korea]
  • Pontoclausia prima (Rocha C.E.F., 1989) (syn. Doviella prima Rocha C.E.F., 1989)
  • Pontoclausia pristina Lee et al., 2022 [Korea]
  • Pontoclausia tomis Bacescu & Por, 1959
  • Pontoclausia wilsoni (Gooding, 1963)

    MESNILLA Canu, 1898

  • Mesnilia cluthae (Scott T. & Scott A., 1896)
  • Mesnilia cluthae (Scott T. & Scott A., 1896)

    MEGACLAUSIA O'Reilly, 1995

  • Megaclausia mirabilis (O'Reilly, 1995)

    INDOCLAUSIA Sebastian and Pillai, 1974

  • Indoclausia bipartita Kim, 2014 [Korea]
  • Indoclausia bacescui Sebastian and Pillai, 1974 [West Indies]
  • Indoclausia indica Sebastian and Pillai, 1974 [West Indies]
  • Indoclausia helioporae Chang & Teng, 2014 [Taiwan]

    SPIONICOLA Biornberg & Radashevsky, 2009

  • Spionicola mystaceus Biornberg & Radashevsky, 2009

    RHODINICOLA Levinsen, 1878

  • Rhodinicola polydorae Bjornborg & Radashervskj, 2011 [Japan] (syn.)
  • Rhodinicola elongata Levinsen 2003 [Korea]
  • Rhodinicola gibbosa Bresciani, 1964
  • Rhodinicola laticauda Ho & Kim, 2003 [Korea]
  • Rhodinicola polydorae Björnberg & Radashevsky, 2011
  • Rhodinicola rugosa Bresciani, 1964 (syn. Rhodinicola rugosa (Giesbrecht, 1897)
  • Rhodinicola similis Kim I.H., Sikorski, O'Reilly & Boxshall, 2013
  • Rhodinicola tenuis Kim I.H., Sikorski, O'Reilly & Boxshall, 2013
  • Rhodinicola thomassini Laubier, 1970 [Indian Ocean]

    PSEUDOCLAUSIA Bocquet & Stock, 1960

  • Pseudoclausia giesbrechti Bocquet & Stock, 1960
  • Pseudoclausia longiseta Bocquet & Stock, 1963
  • Pseudoclausia tomisi Bacescu & Por, 1959




    POLYANKYLIIDAE Ho & Him, 1997


    POLYANKYLIS Ho & Him, 1997

  • Polyankylis australis Karanovic, 2008[Australia; marine interstitial]
  • Polyankylis bogilensis Lee et al., 2022 [Korea]
  • Polyankylis ovilaxa Hamond, 2009
  • Polyankylis orientalis Ho & Kim I.H., 1997

    SEWELLOYA Kim I.H., 2014

  • Sewelloya plana Kim I.H., 2014




    CORYCAEIDAE Dana, 1852

    (Corycella Farran, 1911 = Farranula Wilson C.B., 1932; Mimocorycella Rose, 1929 syn. Corycaeus Dana, 1845 (Rose (1933)

    AGETUS Krřyer, 1849 (syn. Corycaeus (Agetus) Krřyer, 1848)

  • Agetus flaccus (Giesbrecht, 1891)
  • Agetus limbatus (Brady, 1883)
  • Agetus typicus Krřyer, 1849

    CORYCAEUS Dana, 1845 (syn. Corycaeus (Corycaeus) Dana, 1845; Mimocorycella Rose, 1929 (Rose (1933)

  • Corycaeus (Onchocorycaeus) agilis Dana, 1849 (syn. Onychocorycaeus agilis (Dana, 1849)
  • Corycaeus (Onchocorycaeus) catus Dahl F., 1894 (syn. Onychocorycaeus catus (Dahl F., 1894)
  • Corycaeus clausi Dahl F., 1894
  • Corycaeus crassiusculus Dana, 1849
  • Corycaeus obtusus Dana, 1849
  • Corycaeus speciosus Dana, 1849
  • Corycaeus vitreus Dana, 1849

    DITRICHOCORYCAEUS Dahl M., 1912 (syn. Corycaeus (Ditrichocorycaeus) Dahl M., 1912

  • Ditrichocorycaeus affinis (McMurrich, 1916)
  • Ditrichocorycaeus africanus (Dahl F., 1894)
  • Ditrichocorycaeus agilis (Dana, 1849) (syn. Onychocorycaeus agilis (Dana, 1849)
  • Ditrichocorycaeus amazonicus (Dahl F., 1894)
  • Ditrichocorycaeus americanus (Wilson M.S., 1949)
  • Ditrichocorycaeus andrewsi (Farran, 1911)
  • Ditrichocorycaeus anglicus (Lubbock, 1857)
  • Ditrichocorycaeus asiaticus (Dahl F., 1894)
  • Ditrichocorycaeus aucklandicus (Krämer, 1895)
  • Ditrichocorycaeus brehmi (Steuer, 1910)
  • Ditrichocorycaeus dahli (Tanaka, 1957)
  • Ditrichocorycaeus dubius (Farran, 1911)
  • Ditrichocorycaeus erythraeus (Cleve, 1904)
  • Ditrichocorycaeus lubbocki (Giesbrecht, 1891)
  • Ditrichocorycaeus minimus (Dahl F., 1894)
  • Ditrichocorycaeus minimus indicus Oljavidjak & Bojanic, 2008 [Adriatic Sea]
  • Ditrichocorycaeus subtilis (Dahl M., 1912)
  • Ditrichocorycaeus tenuis (Giesbrecht, 1891)

    ONYCHOCORYCAEUS M. Dahl, 1912

  • Onychocorycaeus pacificus (Dahl, 1894)
  • Onychocorycaeus catus (Dahl, 1894)

    FARRANULA Wilson C.B., 1932 (syn. Corycaeus (Corycella) Farran, 1911; Corycella Farran, 1911)

  • Farranula brevis (Farran, 1911) (syn. Farranula gibbula (Giesbrecht, 1891)
  • Farranula carinata (Giesbrecht, 1891)
  • Farranula concinna (Dana, 1849)
  • Farranula curta (Farran, 1911)
  • Farranula dahlae Wi, Soh, Jeong & Kang, 2015
  • Farranula gibbula (Giesbrecht, 1891)
  • Farranula gracilis (Dana, 1849)
  • Farranula longicaudis (Dana, 1849)
  • Farranula orbisa Wi & Soh, 2013
  • Farranula rostrata (Claus, 1863)

    UROCORYCAEUS Dahl, 1912 [syn. Corycaeus (Urocorycaeus) Dahl M., 1912)]

  • Urocorycaeus furcifer (Claus, 1863)
  • Urocorycaeus lautus (Dana, 1849)
  • Urocorycaeus longistylis (Dana, 1849)

    MONOCORYCAEUS Dahl M., 1912 [syn.Corycaeus (Monocorycaeus) Dahl M., 1912]

  • Monocorycaeus robustus (Giesbrecht, 1891)




    SCHMINKEPINELLIDAE Martinez Arbizu, 2006

    "Cyclopoids of the family Schminkepinellidae were discovered in the deep-sea and in marine caves (Martínez Arbizu 2006). The genera initially allocated to Schminkepinellidae were the monotypic genera Cyclopinella G.O. Sars, 1913 Barathricola Humes, 1999, Einslepinella Martínez Arbizu, 2006, Muceddina Jaume & Boxshall, 1996, and Schminkepinella Martínez Arbizu, 2006. The family was considered as the sister group of Poecilostomatoida allocated to the order Cyclopoida (Martínez Arbizu 2000) (from Ivanenko et al., 2019)


    SCHMINKEPINELLA Martinez Arbizu, 2006

    Type-species: Schminkepinella plumifera Martinez Arbizu. 2006

  • Schminkepinella plumifera Martinez Arbizu, 2006


    EINSLEPINELLA Martinez Arbizu, 2006

    Type-species: Einslepinella ulrichi Martinez Arbizu (in press)

  • Einslepinella alignatha Martinez Arbizu, 2006
  • Einslepinella mediana Martinez Arbizu, 2006
  • Einslepinella ulrichi Martinez Arbizu. 2006


    CYCLOPINELLA Sars, 1913

    Type-species: Cyclopinella tumidula Sars, 1913

  • Cyclopinella glacialis (Brady, 1910) [Germany, Scotland; marine] *
  • Cyclopinella multispinosa Jaume & Boxshall,1996 (syn. Muceddina multispinosa Jaume & Boxshall, 1996)
  • Cyclopinella rimensis Humes, 1999 ( syn. Barathricola rimensis Humes, 1999)
  • Cyclopinella tincambayensis Karanovic, 2008 [Australia; marine interstitial]
  • Cyclopinella tumidula Sars, 1913 [Norway; marine]

    * incerta sedis


    BARATHRICOLA Humes, 1999

    Type-species: Barathricola rimensis Humes, 1999

  • Barathricola rimensis Humes, 1999 [northeastern Pacific; hydrotermal vents] (syn. Barathricola rimensis Humes, 1999)
  • Barathricola thermophilus Ivanenko et al., 2019 [Indian Ocean]


    MUCEDDINA Jaume & Boxshall, 1996

    Type-species: Muceddina multispinosa Jaume & Boxshall, 1996

  • Muceddina multispinosa Jaume & Boxshall, 1996 [Western Mediterranean; anchialine caves]
  • Muceddina trispinosa Jaume & Boxshall, 1996(= Muceddina multispinosa Jaume & Boxshall, 1996)



    PHYLOGENETIC RELATIONSHIPS WITHIN SCHMINKEPINELLIDAE (from: Martinez-Arbizu, 2006)


    MACROCHIRONIDAE Humes & Boxshall, 1996

    The family Macrochironidae is composed of five genera: Macrochiron Brady, 1872, Paramacrochiron Reddiah, 1969, Pseudomacrochiron Reddiah, 1969, Oncaeola Kramer, 1895 and Sewellochiron Humes, 1969. All genera are associated with marine invertebrates,

    MACROCHIRON Brady, 1872 (syn. Macrocheiron Brady, 1872 )

    The genus Macrochiron Brady, 1872 currently comprises 24 species, most of which live in association with the hydroid genera Aglaophenia, Lytocarpus and Gymnangium.

  • Macrochiron angulare Humes, 1977
  • Macrochiron anomalum Kim I.H., 2000
  • Macrochiron avirostrum Morris, 1973
  • Macrochiron buddhensis (Thompson I.C. & Scott A, 1903) (syn. Critomolgus buddhensis (Thompson I.C. & Scott A., 1903)
  • Macrochiron brevipes Shen & Lee, 1966 (syn.Critomolgus brevipes (Shen & Lee, 1966)
  • Macrochiron cheliferum (Thompson I.C. & Scott A., 1903)
  • Macrochiron echinicolum Humes & Stock, 1973
  • Macrochiron elegans (Thompson & Scott A., 1903) (syn. Lichomolgus elegans Thompson I.C. & Scott A., 1903)
  • Macrochiron fucicolum Brady, 1872
  • Macrochiron gracilipes (Scott A., 1909) (syn. Doridicola gracilipes (Scott A., 1909)
  • Macrochiron hudsoni Morris, 1973
  • Macrochiron hirsutipes (Scott T., 1893) (syn. Doridicola hirsutipes (Scott T., 1893)
  • Macrochiron lobatum Humes & de Maria, 1969
  • Macrochiron lytocarpi Humes, 1966
  • Macrochiron malayense Sewell, 1949 (syn. Pseudomacrochiron malayense (Sewell, 1949)
  • Macrochiron mutatum Stock, 1957
  • Macrochiron ornatus Krishnaswamy, 1952 (syn. Pseudomacrochiron ornatum (Krishnaswamy, 1952)
  • Macrochiron parvum (Scott A., 1909) (syn. Pseudomacrochiron parvum (Scott A., 1909)
  • Macrochiron rigida Ummerkutty, 1962 (syn. Critomolgus rigidus (Ummerkutty, 1962)
  • Macrochiron rostratum Humes, 1966
  • Macrochiron sargassi Sars G.O., 1916
  • Macrochiron spinipes Sewell, 1949 (syn.Anisomolgus spinipes (Sewell, 1949)
  • Macrochiron trochi (Canu, 1899) (syn. Epimolgus trochi (Canu, 1899) (synonym)
  • Macrochiron valgum Humes, 1966
  • Macrochiron vervoorti Humes & de Maria, 1969

    KEYS TO THE SPECIES IN THE GENUS Macrochiron


    Subfamily Macrochironinae Sewell, 1949

    PARAMACROCHIRON Sewell, 1949

  • Paramacrochiron amboinense Mulyadi, 2005
  • Paramacrochiron chironecicola Ohtsuka, Metillo & Boxshall, 2015
  • Paramacrochiron ennorense Reddiah, 1968
  • Paramacrochiron japonicum Humes, 1970
  • Paramacrochiron maximum (Thompson I.C. & Scott A., 1903)
  • Paramacrochiron pacificum (Wilson C.B., 1950)
  • Paramacrochiron rhizostomae Reddiah, 1968
  • Paramacrochiron sewelli Reddiah, 1968
  • Paramacrochiron thailandicum Ohtsua et al, 2012 [Thailand]
  • Paramacrochiron tridentatum Ohtsuka, Ha & Thu, 2020 [Vietnam]

    PSEUDOMACROCHIRON Reddiah, 1969

  • Pseudomacrochiron aureliae Tang et al., 2011 [Japan]
  • Pseudomacrochiron fucicolum (Brady, 1872) (syn. Macrochiron fucicolum Brady, 1872)
  • Pseudomacrochiron malayense (Sewell, 1949)
  • Pseudomacrochiron ornatum (Krishnaswamy, 1952)
  • Pseudomacrochiron parvum (Scott A., 1909)
  • Pseudomacrochiron pocilloporae Kim I.H., 2004
  • Pseudomacrochiron stocki Reddiah, 1969
  • Pseudomacrochiron urostenum Kim I.H., 2000

    SEWELLOCHIRON Humes, 1969
  • Sewellochiron fidens Humes, 1969

    ONCAEOLA Kramer, 1895 (taxon inquirendum, treated as a genus inquirendum by Boxshall & Halsey (2004)




    PRAXILLINICOLIDAE Huys, 2016

    PRAXILLINICOLA M'Intosh, 1885

  • Praxillinicola kroyeri M'Intosh, 1885





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