FAMILY MIRACIDAE Dana, 1846

Subfamily Diosaccinae Sars, 1906


The Miracidae (ex Diosaccidae Sars, 1906) is a predominantly marine family, but containing several freshwater species, most of them in the genus Schizopera.

The family presently includes 479 species and subspecies, three subfamilies and 43 valid genera [DIOSACCINAE: Actopsyllus, Amonardia, Amphiascus, Amphiascopsis, Antiboreodiosaccus, Balucopsylla, Bulbamphiascus, Dactylopodamphiascopsis, Diosaccus, Diosaccopsis, Eoschizopera, Haloschizopera, Helmutkunzia, Ialysus, Metamphiascopsis, Miscegenus, Neomiscegenus, Paradiosaccus, Paramphiascella, Paramphiascoides, Paramphiascopsis, Pararobertsonia, Parialysus, Pholenota, Protopsammotopa, Psammotopa, Pseudamphiascopsis, Pseudodiosaccopsis, Pseudodiosaccus, Rhyncholagena, Robertgurneya, Robertsonia, Schizopera, Schizoperoides, Schizoperopsis, Sinamphiascus, Teissierella, Tydemanella, Typhlamphiascus; STENHELIINAE: Stenhelia, Pseudostenhelia, Cladorostrata, Onychostenhelia, Melima, Anisostenhelia, Beatricella, Hicksia ; MIRACIINAE: Miracia, Oculosetella, Macrosetella, Distioculus ] with representatives for the most part living in sea waters, a few (Stenhelia, Schizopera, Eoschizopera, Schizoperoides, Cladorostrata, Paramphiascella, Amphiascus) including species living also in fresh or brackish waters, as well as in ground waters (Willen, 2000).

Key to genera in : Lang K. 1965. Copepoda Harpacticoidea from The Californan Pacific coast. K. svenska vetensk. Akad. Handl., 10 (2): 1-566



  • DIOSACCINAE Sars, 1906

  • STENHELIINAE Brady, 1880

  • MIRACIINAE Dana, 1846



    DIOSACCINAE Sars, 1906


    Actopsyllus Wells, 1967

    Amonardia Lang, 1944

    Amphiascoides Lotufo & Fleeger, 1995

    Amphiascopsis Gurney, 1927

    Amphiascus Sars, 1905

    Antiboreodiosaccus Lang, 1944

    Balucopsylla Rao, 1972

    Bulbamphiascus Lang, 1944

    Dactylopodamphiascopsis Lang, 1944

    Dinetocaris Gomez et al., 2021 (syn. Dinetia Gomez et al., 2021)

    Diosaccus Boeck, 1872

    Diosaccopsis Monard, 1928

    Distiocolus Boxshall, 1979

    Eoschizopera Wells & Rao, 1976

    Glabrotelson Huys, 2009

    Goffinella Wilson, 1932

    Haloschizopera Lang, 1948

    Helmutkunzia Wells & Rao, 1976

    Ialysus Brian, 1927

    Macrosetella Sewell, 1940

    Metamphiascopsis Lang, 1944

    Miscegenus Wells, Hicks & Coull, 1982

    Miracia Boxshall, 1979

    Monardius Huys, 2009

    Neomiscegenus Karanovic & Ranga Reddy, 2004

    Pallarica Gomez et al., 2021

    Parialysus Nicholls, 1941

    Paramphiascella Lang, 1948

    Paramphiascoides Wells, 1967

    Paramphiascopsis Lang, 1944

    Pararobertsonia Lang, 1948

    Pholenota Vervoort, 1964

    Protopsammotopa Geddes, 1968

    Psammotopa Pennak, 1942

    Pseudamphiascopsis Lang, 1944

    Pseudobalucopsylla Mu & Huys, 2021

    Pseudodiosaccopsis Lang, 1944

    Pseudodiosaccus T. Scott, 1906

    Rhyncolagena Lang, 1944

    Robertgurneya Lang, 1944

    Robertsonia Brady, 1880

    Sarsamphiascus Huys, 2009

    Schizopera Sars, 1905 s.l.

    Schizoperoides Por, 1968

    Schizoperopsis Apostolov, 1982

    Sinamphiascus Mu & Gee, 2000

    Spinopedia Gomez et al., 2021

    Spinodiosaccus Gomez et al., 2021

    Teissierella Monard, 1935

    Tydemanella A. T. Scott, 1909

    Typhlamphiascus Lang, 1944



    GLABROTELSON Huys in Kihara & Huys (2009) (syn. Hastigerella Nicholls, 1935)

  • Glabrotelson abbotti (Lang, 1965) Huys, 2009
  • Glabrotelson antarcticum (Dahms & Schminke, 1992) Huys, 2009
  • Glabrotelson bengalense (Rao, 1972) Huys, 2009
  • Glabrotelson bodini (Apostolov, 1974) Huys, 2009
  • Glabrotelson bozici (Soyer, 1974) Huys, 2009
  • Glabrotelson chappuisi (Soyer, 1974) Huys, 2009
  • Glabrotelson clavatum (Rao & Ganapati, 1969) Huys, 2009
  • Glabrotelson leptoderma (Klie, 1929) Huys, 2009
  • Glabrotelson mehuinensis (Mielke, 1986) Huys, 2009
  • Glabrotelson meridionale (Chappuis, 1954) Huys, 2009
  • Glabrotelson monniotae (Guille & Soyer, 1966) Huys, 2009
  • Glabrotelson noodti (Rao & Ganapati, 1969) Huys, 2009
  • Glabrotelson psammae (Noodt, 1955) Huys, 2009
  • Glabrotelson scheibeli (Mielke, 1975) Huys, 2009
  • Glabrotelson setosum (Rao & Ganapati, 1969) Huys, 2009
  • Glabrotelson soyeri (Bodin, 1979) Huys, 2009
  • Glabrotelson tenuissima (Klie, 1929)


    SARSAMPHIASCUS Huys, 2009 (syn. Mesamphiascus Nicholls, 1941)

  • Sarsamphiascus amblyops (Sars G.O., 1911)
  • Sarsamphiascus ampullifer (Humes, 1953)
  • Sarsamphiascus angustipes (Gurney, 1927)
  • Sarsamphiascus brevis (Sars G.O., 1909)
  • Sarsamphiascus caudaespinosus (Brian, 1927)
  • Sarsamphiascus congener (Sars G.O., 1909)
  • Sarsamphiascus demersus (Nicholls, 1939)
  • Sarsamphiascus dentiformis (Coull, 1971)
  • Sarsamphiascus discrepans (Mielke, 1989)
  • Sarsamphiascus elongatus (Itô Tat, 1972)
  • Sarsamphiascus gauthieri (Monard, 1936)
  • Sarsamphiascus gracilis (Lang, 1936)
  • Sarsamphiascus graciloides (Klie, 1950)
  • Sarsamphiascus hawaiiensis Yeom & Lee, 2020 [Korea]
  • Sarsamphiascus jackiei Karanovic, 2020
  • Sarsamphiascus jermainei Karanovic, 2020
  • Sarsamphiascus kawamurai (Ueda & Nagai, 2005)
  • Sarsamphiascus kareliensis Apostolov, 2021 (White Sea) (Syn. Amphiascus minutus (Claus, 1863) - sensu Kornev & Chertoprud 2008)
  • Sarsamphiascus hirtus (Gurney, 1927)
  • Sarsamphiascus lobatus (Hicks, 1971)
  • Sarsamphiascus longiarticulatus (Marcus, 1974)
  • Sarsamphiascus michaeli Karanovic, 2020
  • Sarsamphiascus minutus (Claus, 1863) [Germany]
  • Sarsamphiascus pacificus (Sars G.O., 1905)
  • Sarsamphiascus marloni Karanovic, 2020
  • Sarsamphiascus paracaudaespinosus (Roe, 1958)
  • Sarsamphiascus parvus (Sars G.O, 1906)
  • Sarsamphiascus polaris (Sars G.O., 1909)
  • Sarsamphiascus profundus (Becker & Schriever, 1979)
  • Sarsamphiascus propinquus (Sars G.O., 1906)
  • Sarsamphiascus sinuatus (Sars G.O., 1906)
  • Sarsamphiascus tainui (Hicks, 1989)
  • Sarsamphiascus tenellus (Sars G.O., 1906)
  • Sarsamphiascus tenax (Brian, 1927) accepted as Robertgurneya tenax (Brian, 1927)
  • Sarsamphiascus tenuiremis (Brady & Robertson, 1876)
  • Sarsamphiascus titoi Karanovic, 2020
  • Sarsamphiascus ultimus (Monard, 1928)
  • Sarsamphiascus undosus (Lang, 1965)
  • Sarsamphiascus varians (Norman & Scott T., 1905)

    Key to species of Amphiascus Sars, 1905c (modified after Hicks (1986b)


    AMPHIASCUS Sars, 1905 (syn. Paramphiascopsis Lang, 1944)

  • Amphiascus abyssi (Boeck, 1873) accepted as Pararobertsonia abyssi (Boeck, 1873)
  • Amphiascus aegypticus Gurney, 1927 accepted as Amphiascopsis aegypticus (Gurney, 1927) accepted as Sarsamphiascus minutus (Claus, 1863)
  • Amphiascus affinis Sars G.O., 1906
  • Amphiascus aiigrapequensis Pesta, 1959
  • Amphiascus amblyops Sars, 1911 [Norway; marine] (Mesamphiascus amblyops (Sars G.O., 1911) accepted as Sarsamphiascus amblyops (Sars G.O., 1911)
  • Amphiascus ampullifer (Humes, 1953) [north Europe; parasitic] (=Sarsamphiascus ampullifer (Humes, 1953)
  • Amphiascus angolensis Monard, 1934 accepted as Robertsonia angolensis (Monard, 1934)
  • Amphiascus angustipes Gurney, 1927 [Egypt, Italy, Bermuda, France, Sri Lanka; brackish waters] (=Sarsamphiascus angustipes (Gurney, 1927)
  • Amphiascus angrapequensis Pesta, 1916 accepted as Amphiascopsis angrapequensis (Pesta, 1916)


  • Amphiascus attenuatus Sars G.O., 1906 accepted as Amphiascopsis attenuatus (Sars G.O., 1906) accepted as Pseudamphiascopsis attenuatus (Sars G.O., 1906)
  • Amphiascus blanchardi (Scott T. & Scott A., 1895) accepted as Mesamphiascus blanchardi (Scott T. & Scott A., 1895) accepted as Typhlamphiascus blanchardi (Scott T. & Scott A., 1895)
  • Amphiascus banyulensis Monard, 1928 accepted as Amphiascopsis banyulensis (Monard, 1928) accepted as Metamphiascopsis banyulensis (Monard, 1928)
  • Amphiascus brevicornis (Thompson I.C. & Scott A., 1903) accepted as Typhlamphiascus brevicornis Thompson I.C. & Scott A., 1903
  • Amphiascus brunneus Willey, 1932 accepted as Pseudodiosaccopsis brunneus (Willey, 1932)
  • Amphiascus brevis Sars, 1909 [Groenland; marine] (syn. Sarsamphiascus brevis (Sars G.O., 1909)
  • Amphiascus bulbifer Sars G.O., 1911 accepted as Mesamphiascus bulbifer (Sars G.O., 1911) accepted as Haloschizopera bulbifer (Sars G.O., 1911)
  • Amphiascus bulbiseta (Apostolov, 2011)
  • Amphiascus calcarifer Sewell, 1940 accepted as Paramphiascella calcarifer (Sewell, 1940)
  • Amphiascus coralicola Sewell, 1940 accepted as Amphiascopsis coralicola (Sewell, 1940)
  • Amphiascus catharinae Scott T., 1906 accepted as Sarsamphiascus minutus (Claus, 1863)
  • Amphiascus ceylonicus (Thompson I.C. & Scott A., 1903) accepted as Amphiascopsis ceylonicus (Thompson I.C. & Scott A., 1903) accepted as Amphiascopsis cinctus (Claus, 1866)
  • Amphiascus cinctus (Claus, 1866) accepted as Amphiascopsis cinctus (Claus, 1866)
  • Amphiascus commensalis Seiwell, 1928 accepted as Amphiascoides commensalis (Seiwell, 1928) accepted as Paramphiascella commensalis (Seiwell, 1928)
  • Amphiascus confusus (Scott T., 1902) accepted as Mesamphiascus confusus (Scott T., 1902) accepted as Typhlamphiascus confusus (Scott T., 1902)
  • Amphiascus congener Sars, 1909 [north Europe; ?] (=Sarsamphiascus congener (Sars G.O., 1909)
  • Amphiascus clandestinus Klie, 1923 accepted as Schizopera (Schizopera) clandestina (Klie, 1923) accepted as Schizopera clandestina (Klie, 1923)
  • Amphiascus caudaespinosus Brian, 1927 [Algerie; psammon] =Sarsamphiascus caudaespinosus (Brian, 1927)
  • Amphiascus coralifera Sewell (?)
  • Amphiascus ctenophorus Monard, 1928 accepted as Amphiascoides ctenophorus (Monard, 1928) accepted as Robertgurneya rostrata (Gurney, 1927)
  • Amphiascus dactylifer Wilson C.B., 1932 accepted as Robertgurneya dactylifer (Wilson C.B., 1932)
  • Amphiascus debilis (Giesbrecht, 1881) accepted as Amphiascoides debilis (Giesbrecht, 1881)
  • Amphiascus debiloides Monard, 1928 accepted as Amphiascoides brevifurca (Czerniavsky, 1868)
  • Amphiascus demersus Nicholls, 1939 accepted as Sarsamphiascus demersus (Nicholls, 1939)
  • Amphiascus dentatus (Thompson I.C. & Scott A., 1903) accepted as Amphiascopsis dentatus (Thompson I.C. & Scott A., 1903) accepted as Diosaccus dentatus (Thompson I.C. & Scott A., 1903)
  • Amphiascus denticulatus (Thompson I.C., 1893) accepted as Mesamphiascus denticulatus (Thompson I.C., 1893) accepted as Bulbamphiascus denticulatus (Thompson I.C., 1893)
  • Amphiascus dentiformis Coull, 1971 [Virgin Islands, meiobenthic] syn. Sarsamphiascus dentiformis (Coull, 1971)
  • Amphiascus dictydiophorus Monard, 1924 accepted as Robertgurneya dictydiophora (Monard, 1924)
  • Amphiascus discrepans Mielke, 1989 [Chile; marine benthos] (syn. Sarsamphiascus discrepans (Mielke, 1989)
  • Amphiascus dubius Jakubisiak, 1933 accepted as Amonardia similis (Claus, 1866)
  • Amphiascus elegans Lang, 1948
  • Amphiascus elongatus Ito, 1972 [Japan, Antarctic (Livingston Island); interstitial, parasitic] (syn. Sarsamphiascus elongatus (Itô Tat, 1972)
  • Amphiascus ekmani (Lang, 1965) Huys, 2009
  • Amphiascus ecaudatus Monard, 1936 accepted as Robertgurneya ecaudata (Monard, 1936)
  • Mesamphiascus erythraeus (Scott A., 1902) accepted as Robertgurneya erythraeus (Scott A., 1902); Robertgurneya erythraea (Scott A., 1902)
  • Amphiascus exiguus Sars G.O., 1906 accepted as Mesamphiascus exiguus (Sars G.O., 1906) accepted as Haloschizopera exigua (Sars G.O., 1906)
  • Amphiascus falklandiensis Lang, 1936 accepted as Robertgurneya falklandiensis (Lang, 1936)
  • Amphiascus fucicolus Scott T., 1912 accepted as Amphiascopsis fucicolus (Scott T., 1912) accepted as Amonardia similis (Claus, 1866)
  • Amphiascus gauthieri Monard, 1936 [Algerie; psammon] (syn. Mesamphiascus gauthieri (Monard, 1936), Sarsamphiascus gauthieri (Monard, 1936)
  • Amphiascus giesbrechti Sars G.O., 1906
  • Amphiascus glacialis Lang, 1948
  • Amphiascus gracilis Lang, 1936 [Kerguelen; psammon] (=Amphiascopsis gracilis (Lang, 1936); Sarsamphiascus gracilis (Lang, 1936)
  • Amphiascus graciloides Klie, 1950 [Ireland, Helgoland; marine] (Syn. Sarsamphiascus graciloides (Klie, 1950)
  • Amphiascus gracilicaudata (Thompson I.C. & Scott A., 1903) accepted as Typhlamphiascus gracilicaudatus (Thompson I.C. & Scott A., 1903)
  • Amphiascus hamiltoni (Thompson I.C. & Scott A., 1903) accepted as Amphiascopsis hamiltoni (Thompson I.C. & Scott A., 1903) ;Diosaccus hamiltoni (Thompson I.C. & Scott A., 1903)
  • Amphiascus hanseni (Brady, 1899 )(syn. Amphiascopsis hanseni (Brady, 1899);Amphiascus cinctus (Claus, 1866); Amphiascopsis cinctus (Claus, 1866)
  • Amphiascus havelocki (Thompson I.C. & Scott A., 1903) accepted as Amphiascopsis havelocki (Thompson I.C. & Scott A., 1903), Amphiascus cinctus (Claus, 1866), Amphiascopsis cinctus (Claus, 1866)
  • Amphiascus herdmani Scott A., 1896 accepted as Pseudamphiascopsis herdmani (Scott A., 1896)
  • Amphiascus hyperboreus (Scott T., 1903) accepted as Amphiascoides hyperboreus (Scott T., 1903)
  • Amphiascus hirsutus Gurney, 1927 [Suez, Arabian Gulf, South Pacific, Brazil; marine] (syn. Metamphiascopsis hirsutus hirsutus (Thompson I.C. & Scott A., 1903)
  • Amphiascus hispidus (Brady, 1880) accepted as Amphiascoides hispidus (Brady, 1880), Paramphiascella hispida (Brady, 1880)
  • Amphiascus ignotus Lang, 1948
  • Amphiascus ilievecensis Monard, 1935 accepted as Robertgurneya ilievecensis (Monard, 1935)
  • Amphiascus imperator Monard, 1928 accepted as Amphiascopsis imperator (Monard, 1928),Amphiascus cinctus (Claus, 1866), Amphiascopsis cinctus (Claus, 1866)
  • Amphiascus imus (Brady, 1872) accepted as Mesamphiascus imus (Brady, 1872) accepted as Bulbamphiascus imus (Brady, 1872)
  • Amphiascus inermis Sewell, 1940 accepted as Bulbamphiascus imus (Brady, 1872)
  • Amphiascus intermedius (Scott T., 1897) accepted as Amphiascoides intermedius (Scott T., 1897)
  • Amphiascus invaginatus Monard, 1926 accepted as Amphiascoides invaginatus (Monard, 1926)
  • Amphiascus junodi Monard, 1935 accepted as Haloschizopera pygmaea (Norman & Scott T., 1905)
  • Amphiascus kawamurai Hueda & Nagai, 2005 [Japan; nori Cultivation tanks (=Sarsamphiascus kawamurai (Ueda & Nagai, 2005)
  • Amphiascus lagenirostris Sars G.O., 1911 accepted as Mesamphiascus lagenirostris (Sars G.O., 1911)
  • Amphiascus lagunaris Grandori, 1925 accepted as Amphiascopsis lagunaris (Grandori, 1925)
  • Amphiascus lamellifer Sars G.O., 1911 accepted as Typhlamphiascus lamellifer (Sars G.O., 1911)
  • Amphiascus langi Monard, 1936 accepted as Amphiascoides langi (Monard, 1936)
  • Amphiascus latifolius Sars G.O., 1909 accepted as Amphiascopsis latifolius (Sars G.O., 1909)
  • Amphiascus latilobus Monard, 1928 accepted as Amphiascopsis thalestroides (Sars G.O., 1911)
  • Amphiascus linearis Sars G.O., 1907 accepted as Amphiascella linearis (Sars G.O., 1907)
  • Amphiascus linearis Sars G.O., 1907 accepted as Amphiascella linearis (Sars G.O., 1907)
  • Amphiascus longiremis Lang, 1948
  • Amphiascus longirostris Monard, 1928 (=Paramphiascopsis longirostris (Claus, 1863)
  • Amphiascus lobatus Hicks, 1971 [new Zealand; coastal waters] (left)
  • Amphiascus longarticulatus Marcus, 1974 [Ireland, Black sea; marine] (syn. Sarsamphiascus longiarticulatus (Marcus, 1974)
  • Amphiascus maldivensis (Wolfenden, 1905) accepted as Amphiascopsis maldivensis (Wolfenden, 1905)
  • Amphiascus mathoi Monard, 1935 accepted as Mesamphiascus mathoi (Monard, 1935)
  • Amphiascus minutus (Claus, 1863) [Palaearctic, marine, psammon] (=Sarsamphiascus minutus (Claus, 1863)
  • Amphiascus monardi Lang, 1934 accepted as Amphiascopsis monardi (Lang, 1935)
  • Amphiascus monodi Monard, 1952 [Africa, Senegal; coastal waters]*
  • Amphiascus mucronatus Lang, 1948
  • Amphiascus nana Sars G.O., 1906 accepted as Amphiascoides nanus (Sars G.O., 1906)
  • Amphiascus nanoides Sars G.O., 1911 accepted as Amphiascoides nanoides (Sars G.O., 1911)
  • Amphiascus nasutus Sars G.O., 1906 accepted as Amphiascopsis nasutus (Sars G.O., 1906)
  • Amphiascus nicobaricus Sewell, 1940 accepted as Metamphiascopsis nicobaricus (Sewell, 1940)
  • Amphiascus normani Sars G.O., 1911 accepted as Mesamphiascus normani (Sars G.O., 1911)
  • Amphiascus obscurus Sars G.O., 1906 accepted as Amphiascus cinctus (Claus, 1866)
  • Amphiascus pacificus Sars, 1905 [Chatham Island; brackish waters, lagoons] (= Mesamphiascus pacificus (Sars G.O., 1905)
  • Amphiascus pallidus Sars G.O., 1906
  • Amphiascus paracaudaespinosus Roe, 1958 [north Europe; coastal waters] syn. Sarsamphiascus paracaudaespinosus (Roe, 1958)
  • Amphiascus paromolae (Soyer, 1973)
  • Amphiascus parvus Sars, 1906 [Europe, Bermuda, north America; marine, psammon] (syn. Mesamphiascus parvus (Sars G.O., 1906)
  • Amphiascus parvulus (Claus, 1866)
  • Amphiascus perplexus (Thompson & A. Scott, 1903) [Sri Lanka, France?; marine]
  • Amphiascus pestai Monard, 1935
  • Amphiascus pyroiedes Monard, 1928
  • Amphiascus phyllopus Sars G.O., 1906 accepted as Amphiascopsis phyllopus (Sars G.O., 1906)
  • Amphiascus polapinquus Marcus, 1966 (syn. Amphiascus polapinquus Marcus, 1966)
  • Amphiascus polaris Sars, 1909 [Groenland;?] (syn. Sarsamphiascus polaris (Sars G.O., 1909)
  • Amphiascus productus Sars G.O., 1906
  • Amphiascus profundus Becker & Schriever, 1979 ( =Sarsamphiascus profundus (Becker & Schriever, 1979)
  • Amphiascus propinquus Sars, 1906 [north Europe; marine] (=Mesamphiascus propinquus (Sars G.O., 1906)
  • Amphiascus proximus Scott T., 1914 accepted as Amphiascoides proximus (Scott T., 1914)
  • Amphiascus rebus Sewell, 1940 accepted as Amphiascopsis rebus (Sewell, 1940)
  • Amphiascus roberti Monard, 1935 accepted as Amphiascoides roberti (Monard, 1935)
  • Amphiascus robinsonii (Scott A., 1902) accepted as Amphiascoides robinsonii (Scott A., 1902)
  • Amphiascus robustus (Thompson I.C. & Scott A., 1903) accepted as Amphiascopsis robustus (Thompson I.C. & Scott A., 1903)
  • Amphiascus rostratus Gurney, 1927 accepted as Robertgurneya rostrata (Gurney, 1927)
  • Amphiascus rufescens Brian, 1925
  • Amphiascus sahelensis Monard, 1936
  • Amphiascus salammboi Klie, 1942
  • Amphiascus scotti Sewell, 1924(syn.Robertsonia propinqua (Scott T., 1894)
  • Amphiascus sexsetatus Monard, 1928 accepted as Amphiascopsis sexsetatus (Monard, 1928)
  • Amphiascus sinuatus Sars, 1906 [north Europe, north America; sea waters]
  • Amphiascus soyeri (Lang, 1965)
  • Amphiascus tainui Hicks, 1989 [New Zealand; sea waters]
  • Amphiascus thalestroides Sars G.O., 1911
  • Amphiascus tenellus Sars, 1906 [Europe, Algerie; psammon]
  • Amphiascus tenuiremis (Brady & Robertson, 1880) [Europe; marine, psammon]
  • Amphiascus triarticulatus (Moore, 1976)
  • Amphiascus ultimus Monard, 1928 [France; marine] *
  • Amphiascus undosus Lang, 1965 [USA; coastal waters]*
  • Amphiascus varians (Norman & Scott, 1905) [north Europe; marine, psammon, amongst algae] *
  • Amphiascus waihonu (Hicks, 1986)

    * questionable species

    Key to species of Amphiascus Sars, 1905c (modified after Hicks (1986b))


    PARAROBERTSONIA Lang, 1948


  • Pararobertsonia abyssi Lang, 1948
  • Pararobertsonia chesapeakensis Lang, 1948


    PSAMMOTOPA Pennak, 1942 (syn.Protoleptastacus Noodt, 1952 )

  • Psammotopa biarticulata Mielke, 1990 [Panama; marine interstitial]
  • Psammotopa chappuisi Noodt, 1955 [France, Spain, north Africa; marine interstitial]
  • Psammotopa phyllosetosa (Noodt, 1952) [Europe; marine interstitial]
  • Psammotopa polyphylla Noodt, 1955 [France, Portugal; marine interstitial]
  • Psammotopa trisetosa Mielke, 1995 [Venezuela; marine interstitial]
  • Psammotopa vulgaris Pennak, 1942 [Europe, Israel, USA; marine interstitial]


    PROTOPSAMMOTOPA Geddes, 1968

  • Protopsammotopa norvegica Geddes, 1968 [Norway; marine interstitial]
  • Protopsammotopa wilsoni Wells, 1977 [USA; marine interstitial]
  • Protopsammotopa tipperi Bouck & Thistle, 2003 [Mexico; marine]


    BALUCOPSYLLA Rao, 1972

  • Balucopsylla similis Rao, 1972 [India; marine interstitial]
  • Balucopsylla triarticulata Wells & Rao, 1987 [Andaman, Nicobar islands, Costa Rica; marine interstitial]


    NEOMISCEGENUS Karanovic & Ranga Reddy, 2004

  • Neomiscegenus indicus Karanovic & Ranga Reddy, 2004 [India; ground waters]


    MONARDIUS Huys, 2009 (syn. Teissierella Monard, 1935)

  • Monardius salammboi (Monard, 1935b) comb. nov. Huys, 2009
  • Monardius massiliensis (Bodin, 1964) comb. nov. Huys, 2009



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